PhD Chapter 3

Results 3/3

This series of files compile all analyses done during Chapter 3:

All analyses have been done with R 4.1.0.

Click on the table of contents in the left margin to assess a specific analysis.
Click on a figure to zoom it

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1. Methodology

The aim of this section is to predict the structure of benthic communities based on the values of environmental variables.

We used abiotic parameters or indices of human exposure indices (calculated in Section 1) as predictors and presence/absence or density data as reponse variables. We tested GLMs and HMSC for now. Each method has been developed in dedicated scripts, and final objects were imported.

First, we computed models using the 108 stations with abiotic variables or exposure indices as predictors. 85 % of the stations (92) act as training data, and the rest (16) will be used to validate the outputs. Second, these models will be used to predict taxa richness and distribution in the entire study area using predictor rasters.

Results are presented with a table regrouping McFadden’s (presence/absence) or Tjur’s pseudo-R2 (density), validation ratios and variables coefficients. Maps display results of the SDM, with the probability of presence of each taxon for which the pseudo-R2 is higher than 0.20 (grey: low probability, dark blue: high). Stations are either plotted with colors (green = taxa present, red = taxon absent) or with circles (wider circles = higher taxon density).

2. Models

2.1. Generalized Linear Models

2.1.1. Presence/absence data

We considered presence/absence data with a binomial distribution. Diagnostics are here.

All parameters
McFadden’s pseudo-R2 and variables coefficients for each taxon
  Pseudo-R2 intercept depth om gravel silt clay arsenic cadmium copper mercury aquaculture dredging runoff sewers structures shipping fisheries
aceroides_aceroides_latipes 0.17 -15.9 -0.124 -0.551 -50.8 0.181 -1.21 -0.787 -0.314 0.768 -0.0623 0.381 -0.586 -0.261 0.418 -0.118 0.274 -0.00927
akanthophoreus_gracilis 0.32 -1.87 0.713 -0.47 -0.107 0.43 0.778 0.535 -1.48 0.706 0.0815 1.14 -0.32 1.1 -0.0226 -1.51 0.807 0.419
ameritella_agilis 0.48 -7.67 0.617 0.372 0.153 -0.868 1.65 -2.84 1.44 1.97 -5.95 0.64 3.7 1.61 2.4 -9.33 -0.415 0.00462
ameroculodes_edwardsi 1 -651 170 -239 75.6 83.9 8.81 -502 28.7 359 -234 -0.509 380 141 -19.2 -541 -301 -143
ampelisca_vadorum 1 -91.5 -12.3 37.4 15.4 1.55 11.4 40.4 -30.5 -78.7 -9.61 -0.145 -13.3 18.3 39.7 -37.9 25.3 14.3
amphipoda 0.12 -0.69 0.458 -0.491 0.201 0.175 -0.106 -0.433 0.488 0.0872 0.538 0.161 0.839 0.885 0.367 -1.57 -0.327 0.0912
anonyx_lilljeborgi 0.36 -383 -1.66 -0.587 1.67 0.322 -2056 -3.52 -0.526 1.79 -0.983 0.388 -4.3 -3.2 -0.0295 1.73 0.354 -4.23
anthozoa 1 -43.1 -7.65 0.529 1.89 16.4 -3.6 -1.28 -13.5 -0.77 -11.3 -8.72 -1.72 -17.8 -1.47 10.2 4.44 -2.5
arcteobia_anticostiensis 1 -65 -1.81 -36.7 9.64 23.2 -3.72 4.28 8.86 -3.25 3.53 21.1 2.04 11.3 -12.7 8.89 5.71 -41.7
arrhoges_occidentalis 1 -65 -1.26 4.33 -6.84 -30.1 -2.48 -11.9 7.01 0.767 -0.967 -25.3 6.81 -21.4 -23.9 5.47 19.4 -26.9
astarte_sp 1 -253 55.4 -39.1 -10.4 3.65 4.49 125 -16.4 -34 -227 -19.5 -62 100 59 -70.6 -85.6 9.25
axinopsida_orbiculata 0.28 -1.9 0.385 0.542 -0.31 -0.258 0.472 -0.0697 0.969 -0.224 -1.73 -0.188 0.942 0.355 -0.336 -0.871 0.264 0.522
axiothella_catenata 1 -57.5 10.7 0.959 -23.7 -12.7 3.02 9.84 2.95 -21.5 43.5 -21.9 -38.8 -15.3 -13 35.5 -16.5 2.44
bathymedon_longimanus 1 -346 -114 -24.3 18.3 46.2 -658 -222 -31 46.3 -40.8 127 94.2 -59.9 212 -30 -3.4 26.7
bathymedon_obtusifrons 1 -323 -51.5 -14.7 -90.6 -0.654 -57.1 18.2 -5.93 72.5 -218 102 -261 -25.8 183 49.1 -44.6 -30.8
bipalponephtys_neotena 0.54 8.52 -1.57 5.27 0.214 0.439 -0.0501 -3.14 0.389 -4.11 8.2 -0.165 0.601 -1.23 -0.776 1.26 4.71 0.386
boreochiton_ruber 1 -222 -28.1 -60.4 26.7 59.7 -202 39 -77.9 5.85 52.4 -19.6 -18.8 -70.9 -57.3 168 -139 4.59
brachydiastylis_sp 1 -106 18.3 22.8 10.9 10.8 16.7 54.6 0.25 -102 7.71 -9.55 -25.8 -0.408 -25.5 36.9 60.4 18.4
byblis_gaimardii -9.95 -2.2e+15 4.84e+13 3.64e+13 -7.76e+13 -3.12e+14 6.24e+13 -2.4e+14 8.32e+13 -8.95e+14 8.43e+14 1.08e+14 4.84e+14 -4.55e+11 -7.02e+14 1.23e+14 -5.02e+13 1.76e+14
cancer_irroratus 1 -117 2.73 -67 39.8 50.2 -2.48 41.3 -0.555 -64.6 3.14 36 22.9 52.4 88 -53.8 6.68 21.8
caprella_septentrionalis -10.38 -1.78e+15 1.02e+14 -7.75e+14 3.74e+13 -6.22e+13 1.26e+14 -7.34e+14 -1.65e+14 9.67e+13 -1.79e+14 6.33e+13 2.43e+14 5.88e+14 8.66e+14 -7.54e+14 -1.07e+15 -1.21e+14
chaetodermatida 0.41 -4.81 2.27 -0.439 -0.913 -1.49 1.73 -7.41 0.893 2.1 -0.561 1.37 0.794 2.47 2.12 -4.37 0.946 0.481
chionoecetes_opilio 1 -138 37.6 4.3 8.1 6.35 27.2 -109 -4.73 60.9 5.03 17.4 -18.2 -38.7 -49.6 29.3 27.2 -110
chlamys_islandica 1 -38.1 -12.8 -2.68 -0.299 5.36 -3.56 0.146 -2.8 0.643 -1.62 -2.37 1.08 -8.36 0.796 1.95 2.5 7.18
chone_sp 1 -120 -8.1 30.2 11.6 -4.43 -2.14 41.9 24.1 -38.9 17.1 58 -105 95 59.4 -27 -1.68 42.8
ciliatocardium_ciliatum 1 -61 4.32 34.1 -14.1 -47.7 -16 -0.189 8.19 -3.47 -1.75 -17.5 -1.01 -41.1 -6.43 25.1 0.0376 2.5
cirripedia 1 -36.2 1.25 1.21 -0.244 -2.37 0.441 -4.44 11.3 -11.6 -1.62 0.275 1.21 -10.8 0.953 13.4 7.69 2.95
cistenides_granulata 0.24 -2.1 -0.304 -0.0374 -0.0344 0.264 -0.827 0.4 -0.72 -1.19 0.427 -0.0419 -0.781 -0.968 -0.28 1.56 -0.169 0.377
cossura_longocirrata 1 -89.2 -27 -6.77 14 33.1 32.8 -20.9 28.5 10.7 1.01 1.05 11.2 32.3 -78.5 -22.4 44.6 -0.862
crassicorophium_bonellii 1 -108 21.9 -38.6 -12.5 -13.8 2.87 16.3 -44 119 -0.239 22 0.268 -11 -68.8 -2.39 51.6 10.5
crenella_decussata 1 -148 25.3 -126 20.8 40.8 -4.17 38.4 -111 -11.7 55.7 3.71 4.23 -27.5 26 16.8 -56.3 -17.9
cumacea 1 -46.4 18.2 -13.6 -13.7 -7.74 4.7 2.59 -16.1 23.4 1.77 4.77 9.22 1.11 15.8 -24.5 -11.1 -8.65
cyclocardia_borealis 1 -43.1 -7.65 0.529 1.89 16.4 -3.6 -1.28 -13.5 -0.77 -11.3 -8.72 -1.72 -17.8 -1.47 10.2 4.44 -2.5
cyrtodaria_siliqua 1 -92.1 24 -39.5 -0.581 -25.9 3.55 -35.8 -29.5 62.7 8.22 47.8 41.3 49.5 41.7 -72.8 4.23 -5.91
diastylis_rathkei 0.4 -347 -1.84 0.0782 0.488 -0.00965 -1865 1.16 -0.191 -0.454 1.25 -2.82 0.48 -0.0133 -4.43 2.51 -1.47 -0.464
diastylis_sculpta 0.65 -180 -4.23 1.04 -79.1 0.473 -824 6.44 -0.276 -3.01 1.31 -1.41 -5.42 -0.554 -8.11 9.03 1.82 -0.12
diastylis_sp 1 -104 32.9 -9.98 17.2 40.6 16.3 -0.292 29.8 5.05 -18.9 -45.6 11.7 -27.7 -22.4 31.4 -31.2 1.04
echinarachnius_parma -8.19 -1.85e+15 2.9e+14 -1.82e+15 -8e+13 5.41e+14 -5.4e+14 -1.11e+15 -3.24e+14 1.09e+15 -9.54e+14 6.05e+14 6.92e+12 -2.18e+14 7.79e+14 -5.68e+13 -1.33e+14 -1.67e+13
edotia_montosa 1 -118 1.61 -8.64 -33.5 -37.7 -9.75 5.36 1.58 37.7 -81.4 -32 -2.32 -57.9 20.5 30.5 -45.2 -6.58
ennucula_tenuis 0.4 -0.658 1.39 -0.135 0.419 0.846 0.812 0.838 -0.89 -1.7 1.02 1.05 -0.523 1.59 0.0536 -0.228 1.14 0.465
eteone_sp 1 -241 -6.71 -127 -9.11 -2.56 -7.78 7.51 -218 68.5 4.23 -44.7 129 -33.7 179 -241 -22.3 -14.5
euchone_sp 1 -352 -59.6 35.7 3.26 -18.1 -10.3 -13.2 18.6 54.2 -244 127 -315 81.5 151 -18.8 2.17 -22.8
eudorella_emarginata 1 -613 -433 71.7 203 287 -91.9 151 -172 -463 3.09 53.4 -58.6 -412 363 128 85.6 35.3
eudorellopsis_integra 0.52 1.84 0.176 1.48 -0.216 0.209 0.398 1.92 0.204 -3.99 4.37 -0.393 -0.57 0.491 -2.4 1.15 3.25 0.171
euspira_pallida 1 -138 16.8 27.6 31.3 56.4 34.1 -55.9 15.3 -16.3 -82.3 -5.06 59.7 86.2 24 -167 49.1 4.02
glycera_capitata 1 -74.3 10.6 -53.5 17.1 13.7 18.3 -64.8 23.2 59.4 -11.3 5.63 15.9 -20 -0.472 -12.1 6.09 -13.7
glycera_sp 1 -792 -263 -77.3 229 84.7 -107 -142 -22.6 111 -19.2 18.4 -655 -289 -47 159 20.6 -716
goniada_maculata 0.49 -290 1.73 0.366 -1.89 0.0107 -1574 -2.9 0.915 -0.48 0.459 0.729 0.248 -0.222 0.894 -0.721 0.621 0.465
guernea_prinassus_nordenskioldi 1 -147 12.6 -131 -27.5 66.6 -49.8 -10.3 -15.2 45.3 -89.6 8.43 -34.6 -11.2 39.2 14.5 3.97 -7.93
halacaridae_spp 1 -38.1 -12.8 -2.68 -0.299 5.36 -3.56 0.146 -2.8 0.643 -1.62 -2.37 1.08 -8.36 0.796 1.95 2.5 7.18
haminoea_solitaria 1 -93.8 -16.7 81.2 -12.3 -57.7 -14.9 24.4 20.4 -38 -8.02 49 -29.4 53.6 10.7 -13.8 27.3 14.7
hardametopa_carinata 1 -352 -59.6 35.7 3.26 -18.1 -10.3 -13.2 18.6 54.2 -244 127 -315 81.5 151 -18.8 2.17 -22.8
harmothoe_sp 1 -386 113 -229 153 387 -41.5 -135 111 132 -222 -163 88.2 -51.1 29.5 -78.3 17.9 -19.3
harpacticoida 0.3 -0.32 -0.658 0.613 -0.0813 -0.814 -0.455 1.35 0.0521 -1.19 -0.202 -0.188 -1.67 2.01 0.0492 -0.847 0.562 0.304
hediste_diversicolor 0.72 -42.6 -20.5 2.67 5.07 9.68 -4.44 3.27 -1.5 -14.3 5.76 9.42 -6.1 -6.27 12.8 23.5 -11.3 -30.8
heteranomia_squamula 1 -60.7 -2.15 -30.8 21.2 33.1 -2.22 27.9 -3.73 -26.5 5.04 6.68 -24.5 13.5 -16.1 29.4 15.9 -13.9
hiatella_arctica 1 -94.2 4.78 -33.3 -3.6 -20.2 12.4 -23.5 -7.92 -7.46 15.6 -10.2 -19.6 -36.9 1.96 49.7 -1.56 -8.27
holothuroidea 1 -77.6 -26.4 29.4 5.3 -13.7 4.88 20.7 -49.7 18.8 -7.06 5.8 -23 45.2 -1.27 -35.5 7.76 11.1
idotea_phosphorea 1 -175 -79.1 102 64.5 35 -61.7 29.6 -62.9 -28.5 -126 56.1 -18.7 171 124 -191 13.1 31.9
ischyroceridae_spp 1 -104 -20.4 76.4 8.34 -13.8 7.06 35.4 21.3 -60.1 11.5 35.2 -54.6 90.7 -24.5 17.2 -1.49 9.6
ischyrocerus_anguipes 1 -355 -129 105 58.5 -80.3 57.4 -400 18.7 130 -67.5 -82.9 -203 -176 -30.1 38.7 51.5 -162
isopoda 1 -41.9 0.299 24.8 -7.04 -20.4 -4.78 8.63 6.46 -13.8 6.44 5.29 -6.3 22.6 -4.91 -10.2 9.21 10.5
lacuna_vincta 1 -196 -125 -71.9 41.2 51.4 -49.2 -108 -33 30.7 -19.8 26.7 31.1 -79.7 75.5 -14.4 28.6 -6.3
lamprops_fuscatus 0.23 -3.17 -0.0931 -1.55 -0.698 0.18 -0.9 -1.57 -0.447 1.18 -0.484 0.964 0.691 -0.219 0.888 -0.183 0.478 0.139
lamprops_quadriplicata 1 -79.8 4.84 4.91 -23.3 -59.3 -19 19.6 5.38 -13.8 11.1 24.3 3.33 -5.8 44.9 1.29 2.79 15.5
lepeta_caeca 1 -621 9.97 -165 47.6 114 -30.3 -682 0.0488 231 -161 -47.1 -312 -155 78.4 41.6 -186 -30.6
leucon_leucon_nasicoides 0.4 -0.711 1.15 2.33 -1.19 -2.23 1.11 0.194 0.513 -1.28 1.35 0.289 -1.09 -1.26 -0.32 1.76 1.1 0.131
littorina_littorea 1 -250 2.54 120 39 -23.5 -23.6 86.8 38.6 -88.9 -94.1 -24.6 -142 168 16.5 12.5 -115 9.4
lumbrineridae_spp 1 -106 28 8.39 15.6 31 13.6 -1.04 43.1 -17.3 -13.9 -51.6 3.26 -47.6 -41.7 67.4 -7.93 6.29
lysianassidae_spp 1 -77.1 -33 7.94 14.4 18.6 -0.0925 0.0502 -35.1 28.3 -29.4 32.5 -6.29 7.79 19.5 -2.49 10.9 10.4
macoma_calcarea 0.23 1.66 -0.146 0.0157 0.171 1.2 -1.35 0.132 -0.269 -1.13 -0.488 -0.0398 0.766 2.09 0.413 -1.99 0.0484 0.276
maera_danae 1 -120 -8.1 30.2 11.6 -4.43 -2.14 41.9 24.1 -38.9 17.1 58 -105 95 59.4 -27 -1.68 42.8
maldane_sarsi 1 -109 30.8 -21.3 21.4 16.1 2.39 14.1 27.4 41.3 -25.5 14.6 -39.5 1.4 23.5 32.1 12.7 28
maldanidae_spp 0.3 -13.4 0.801 0.391 -41.3 0.315 0.474 -0.264 -1.1 1.39 -0.17 0.768 -0.985 -0.169 0.219 0.161 0.825 -0.101
monoculopsis_longicornis 1 -136 -48.4 129 9.02 -25.6 20.2 49.4 17.6 -109 -13.5 18.6 29.2 157 -45 -150 37.7 42.5
muculus_musculus_discors 1 -38.1 -12.8 -2.68 -0.299 5.36 -3.56 0.146 -2.8 0.643 -1.62 -2.37 1.08 -8.36 0.796 1.95 2.5 7.18
mytilus_sp 0.4 -8.72 0.936 0.485 -0.195 0.878 -3.16 -11.8 0.376 2.47 -1.82 0.0602 -0.115 0.0614 2.32 -0.78 -3.62 -0.572
nematoda 0.64 -3.2 2.02 -3.58 -0.504 -0.389 -6.45 3.84 -4.17 0.163 -0.177 1.46 -3.43 1.48 3.29 -2.86 -1.31 -0.551
nemertea 1 -70.1 13.2 -49.5 10.5 4.19 5.29 -67.8 24.3 52 3.46 4.24 24.6 -0.826 -0.971 -31 5.26 -2.91
neoleanira_tetragona 1 -61.7 7.69 5.61 -32.7 -10.8 1.22 26.5 -5.1 -49.6 5.18 -0.7 -6.89 11.2 25.2 -18.1 1.53 0.732
nephtyidae_spp 1 -408 79.9 -116 -214 -8.4 -52.8 115 14.1 -8.94 77.9 38.9 226 300 13.6 -393 -218 53.1
nephtys_caeca 0.4 -328 -2.49 1.79 0.807 -0.676 -1768 3.26 1.51 -2.53 -0.792 -0.115 -1.44 1.41 -3.64 2.36 0.061 0.136
nephtys_incisa 0.36 -2 1.33 1.03 0.223 0.0944 1.05 -1.73 -1.01 0.0608 0.825 0.547 -0.722 0.267 0.312 -0.558 0.875 -0.197
nephtys_sp 1 -297 92.6 -128 92.4 181 71.5 -294 89.9 43.4 -89.5 -25.3 126 22.9 22.1 -241 104 -96.8
nuculana_minuta 0.43 -135 1.43 -2.74 0.773 3.32 -676 -5 -0.107 0.241 -0.806 0.507 -13.9 -4.59 0.587 5.52 2.34 0.415
nymphonidae_spp 1 -60.7 -2.15 -30.8 21.2 33.1 -2.22 27.9 -3.73 -26.5 5.04 6.68 -24.5 13.5 -16.1 29.4 15.9 -13.9
oenopota_sp 0.6 -356 -0.257 -2.4 1.36 6.16 -1400 14.5 -1.27 -10.2 3.96 -261 -6.35 3.45 -5.06 7.68 -0.691 1.56
oligochaeta 0.34 -184 -1.2 1.15 -56.5 0.571 -461 -3.14 0.334 0.68 -0.51 -221 0.45 1.1 1.01 -1.94 1.51 0.473
ophelia_limacina 0.57 -147 3.94 1.26 2.95 -1.22 -0.418 -13.7 2.47 6.86 -1.89 -356 -13 -3.6 -5.32 1.94 8.51 -4.99
opheliidae_spp 1 -167 34.5 46.2 18 3.56 8.55 30.8 41.8 -13.9 -95.4 -59.6 -30.7 -12.9 -46.3 59.7 -19.1 17.8
ophiopholis_aculeata 1 -109 -19.4 18.4 10.4 27.3 -190 18.8 -30.9 -12.4 -33.2 24.8 -16 17.3 14.1 -10.4 27.6 20.5
ophiura_robusta 1 -218 47.7 -150 -19.9 32.9 -78.1 137 -156 -53.5 65.3 -2.41 -30.1 -16.9 37.1 34.9 -69.9 1.37
orchomenella_minuta 1 -125 -23.8 61.9 -6.55 -46.9 -3.84 56.3 -167 -0.199 37.1 -22.7 -74.4 12.4 90 -62.4 -17.9 2.74
ostracoda 0.22 -1.59 0.263 0.215 -0.624 0.179 0.908 -2.78 -0.26 1.27 -0.49 0.24 -0.0721 0.69 -0.414 -1.47 1.15 0.0361
pagurus_sp 1 -65 -1.81 -36.7 9.64 23.2 -3.72 4.28 8.86 -3.25 3.53 21.1 2.04 11.3 -12.7 8.89 5.71 -41.7
pagurus_pubescens 1 -108 10.3 -51.1 16.3 -3.63 20.8 -13.1 -20.9 2.95 52.4 -6.27 -11.9 -23.8 -38.2 65.5 -47.9 -40.5
pandalus_montagui 1 -44.9 2.07 -20 6.39 16.2 -2.72 0.0807 -1.41 5.06 11.5 -1.79 7.73 -8.57 -4.36 15.9 -14.5 -0.169
parathyasira_equalis 1 -93.8 -16.7 81.2 -12.3 -57.7 -14.9 24.4 20.4 -38 -8.02 49 -29.4 53.6 10.7 -13.8 27.3 14.7
parvicardium_pinnulatum 1 -656 4.57 -268 63.4 142 -9.22 302 41.7 41.1 -376 58.5 -804 145 -178 293 -77.6 -27.1
periploma_leanum -7.18 -2.95e+15 6.26e+14 -1.84e+13 -3.16e+14 -2.17e+14 -3.92e+14 4.19e+14 1.33e+13 -3.64e+14 1.72e+14 -2.31e+14 -8.01e+14 -5.34e+14 2.51e+14 4.24e+14 5.97e+14 7.68e+13
philine_lima 1 -65 -1.81 -36.7 9.64 23.2 -3.72 4.28 8.86 -3.25 3.53 21.1 2.04 11.3 -12.7 8.89 5.71 -41.7
philomedes_sp 1 -208 90.9 -36.4 9.68 -12.6 26 54.4 13.4 -124 7.07 -39.9 81.6 40.6 18 -78.3 7.38 5.24
pholoe_longa 1 -352 -59.6 35.7 3.26 -18.1 -10.3 -13.2 18.6 54.2 -244 127 -315 81.5 151 -18.8 2.17 -22.8
pholoe_sp 0.14 -0.0233 -0.371 0.896 -0.0339 0.134 0.321 -0.176 -0.457 -0.013 -0.052 -0.0459 -0.658 -0.101 -1.32 0.83 0.606 -0.343
phoxocephalus_holbolli 0.18 -1.7 -0.334 0.0646 -0.147 -0.604 0.00654 0.0561 0.367 -0.305 0.449 -0.163 -0.236 0.326 -0.298 0.203 -0.947 0.0602
polynoidae_spp 0.25 -370 0.21 0.768 -0.139 0.213 -2011 -1.72 -0.831 0.291 -0.0455 -0.372 -0.766 -0.234 0.996 -0.914 -0.297 -0.756
pontogeneia_inermis 1 -153 12.2 -59 61.3 48.6 -3.57 -88.2 26 112 -93 23.4 -53.8 -30.1 -44.6 23.9 47 -111
pontoporeia_femorata 0.44 -82.8 0.513 -0.776 0.148 1.25 -1.77 0.112 0.431 -0.4 2.36 -225 -0.295 -0.55 -0.0415 1.6 -1.27 -0.491
praxillella_praetermissa 0.67 -35.2 2.46 -1.03 -23.5 -2.31 -3 -4.67 4.83 3.46 -4.72 13.2 -0.0356 -24.1 6.97 23.7 6.79 -26.2
propebela_turricula 1 -1888 503 22.5 -586 -464 -4188 887 -624 306 235 48.3 -445 -75.2 58.5 -599 25.7 -95.6
protomedeia_fasciata 0.37 -267 0.0937 0.543 0.219 0.745 -1441 -0.786 1.78 -1.04 1.5 -0.888 0.592 0.773 -2.67 1.02 -0.0856 0.669
protomedeia_grandimana 0.21 -0.367 0.816 0.738 0.187 -0.166 0.0462 1.01 -0.578 -0.703 1 0.222 -0.594 1.17 0.562 -0.364 -0.336 -0.171
puncturella_noachina 1 -115 3.77 -53.9 35 74.5 -7.24 -178 -8.87 117 -32.3 -3.88 39.3 -44.5 -9.64 -52.6 26.2 -34.9
quasimelita_formosa 0.23 -1.8 0.732 -0.0751 0.237 0.805 -1.7 -0.126 -0.289 0.263 -0.869 1.3 -0.132 1.58 1.51 -1.53 0.043 0.354
quasimelita_quadrispinosa 1 -104 -20.4 76.4 8.34 -13.8 7.06 35.4 21.3 -60.1 11.5 35.2 -54.6 90.7 -24.5 17.2 -1.49 9.6
retusa_obtusa -2.25 -2.51e+15 -6.93e+14 1.7e+14 5.98e+14 9.1e+14 9.67e+13 6e+14 1.26e+15 -4.32e+14 -2.17e+14 -3.38e+14 -5.27e+14 6.17e+14 -2.02e+15 2e+15 4.18e+14 6.51e+14
sabellidae_spp 0.4 -19.7 -1.56 -0.691 -57.6 -0.192 0.0166 -0.253 0.366 0.0201 0.157 -1.53 -1.26 -3.64 0.0692 3.22 -0.202 -0.326
scoletoma_fragilis 1 -686 -210 -80.1 176 756 302 -335 -84.7 -13.6 -184 -104 -229 -711 -297 992 96.5 -479
scoletoma_sp 1 -57.6 -6.99 -29.5 -1.98 10.9 -0.00422 -22 -2.15 2.32 -5.08 -2.74 33.7 -5.31 7.99 -21 4.78 5.49
scoloplos_sp 1 -120 49 -26.6 -14.4 -39 -9.57 19.6 -23.1 43.8 42.2 55.8 26.1 60.2 68.9 -52.1 -63.3 29.3
serripes_groenlandicus 1 -181 -16.7 -3.84 41.7 77.7 -3.9 -191 8.68 126 -107 21.9 74.2 -181 55.4 1.93 43.8 -21.7
sipuncula 0.29 -14.7 0.23 0.475 -45 1.07 0.69 1.79 -0.079 -1.45 0.0391 0.041 -1.23 0.328 -0.654 1.34 0.439 -0.0891
solamen_glandula 1 -43.1 -7.65 0.529 1.89 16.4 -3.6 -1.28 -13.5 -0.77 -11.3 -8.72 -1.72 -17.8 -1.47 10.2 4.44 -2.5
solariella_sp 1 -359 107 -161 34.8 165 -165 283 -6.89 -340 62.2 4.24 -166 64.6 -83.8 207 40.7 64.7
strongylocentrotus_sp 1 -74.8 -23.1 26.6 14.7 33.8 23.8 27.5 -17.1 -86.3 5.32 -20 -2.68 -10.3 -2.32 4.25 19.7 14.2
tachyrhynchus_erosus 1 -120 49 -26.6 -14.4 -39 -9.57 19.6 -23.1 43.8 42.2 55.8 26.1 60.2 68.9 -52.1 -63.3 29.3
thracia_septentrionalis 0.25 -2.4 -1.91 -0.0932 0.481 0.886 -0.147 -2.18 -0.281 -0.106 -1.07 -0.0139 0.548 -0.821 0.485 -0.53 0.954 -0.0481
thyasira_gouldi 0.32 -1.94 1.07 0.148 0.0096 0.249 -0.448 0.413 -1.97 0.268 0.125 0.859 -1.68 0.887 0.964 -0.882 0.723 0.24
thyasira_sp 1 -120 49 -26.6 -14.4 -39 -9.57 19.6 -23.1 43.8 42.2 55.8 26.1 60.2 68.9 -52.1 -63.3 29.3
trichotropis_bicarinata 1 -104 38.5 -1.8 3.17 -13.3 11.2 -48 37.7 -16.1 -7.95 0.634 50.1 24.5 -2.29 -56.1 37.6 -28.7
turritellopsis_stimpsoni -12.63 -1.77e+15 2.53e+14 6.31e+14 -2.91e+14 -1.43e+15 -1.84e+14 3.55e+14 1.19e+14 6.02e+13 4.55e+14 8.6e+13 -4.8e+14 9.13e+14 -8.39e+14 -3.51e+14 2.86e+14 -1.07e+14
yoldia_myalis 1 -480 -12 35.9 127 -75.2 20.3 10.2 30.3 36.4 15.3 125 -629 -170 42.4 199 15.2 -340
Abiotic parameters
McFadden’s pseudo-R2 and variables coefficients for each taxon
  Pseudo-R2 intercept depth om gravel silt clay arsenic cadmium copper mercury
aceroides_aceroides_latipes 0.12 -15.5 -0.0156 -0.831 -50 0.441 -1.21 -0.279 -0.0706 0.338 0.22
akanthophoreus_gracilis 0.19 -1.46 0.725 -0.642 -0.136 0.525 0.656 0.745 -0.433 -1.2 0.503
ameritella_agilis 0.22 -2.99 -0.0402 1.34 0.0994 -1.26 0.815 0.121 0.841 -0.584 -1.23
ameroculodes_edwardsi 0.34 -281 1.18 -0.0907 0.489 0.515 -1507 -2.69 0.176 -1.23 0.392
ampelisca_vadorum 1 -129 13.8 -14 11.7 0.891 21.6 63.7 -60.4 -68.9 43.3
amphipoda 0.06 -0.634 0.252 -0.222 0.132 0.0419 -0.118 -0.0835 0.461 -0.387 0.552
anonyx_lilljeborgi 0.14 -364 -1.03 -0.383 0.307 0.561 -1973 -0.112 -0.508 -0.283 -0.0611
anthozoa 1 -77.8 -2.56 -29.7 9.63 43 -5.21 13.2 -55.7 24.6 -9.86
arcteobia_anticostiensis 1 -150 -9.76 4.93 1.68 40.3 -16.3 -135 -3.05 -39.9 20.9
arrhoges_occidentalis 1 -246 37.7 -225 -14.1 16.6 24.1 121 -17.6 -89.5 45
astarte_sp 0.36 -247 1.7 -2.8 -0.36 1.05 -1303 -1.24 -2.17 -0.435 -0.323
axinopsida_orbiculata 0.19 -1.79 0.525 0.247 -0.227 -0.219 0.313 0.356 0.727 -0.281 -1.76
axiothella_catenata 1 -89.9 33.6 -14.9 -14.8 -4.52 18.9 -68.4 22.8 0.759 37.1
bathymedon_longimanus 0.34 -266 -0.885 0.864 -1.2 1.05 -1411 -7.14 0.202 -0.196 -0.956
bathymedon_obtusifrons 0.62 -30.4 -1.03 -11.8 -27.8 8.25 -45.3 -3.51 1.69 7.43 -4.68
bipalponephtys_neotena 0.33 3.27 -0.0396 2.37 0.0539 0.396 -0.0344 -1.16 -0.155 -0.804 2.24
boreochiton_ruber 0.67 -331 -1.66 -10 1.17 6.16 -1733 1.45 -5.15 -3.19 1.91
brachydiastylis_sp 1 -302 114 -176 14.4 79.1 88.6 82 -127 -62 53.8
byblis_gaimardii 0.39 -221 1.78 -1.45 -0.00689 -0.755 -1158 0.123 -1.04 -4.44 1.68
cancer_irroratus 1 -197 -71.2 -121 33.5 69.6 -14.1 102 -20.2 -98.6 33.8
caprella_septentrionalis 0.59 -15.2 -10.8 0.821 1.35 -0.841 1.12 -4.45 -0.454 1.16 0.371
chaetodermatida 0.29 -2.67 1.34 -0.659 -1.02 -0.834 0.802 -1.66 0.198 0.326 0.437
chionoecetes_opilio 1 -1624 667 -116 54.1 147 301 274 -943 -111 175
chlamys_islandica 1 -157 -23.2 -123 -11.4 54.9 -74 32.9 -60 -3.26 21.2
chone_sp 1 -1544 -951 125 -544 -45.2 -10.2 -502 -1.57 162 -59.1
ciliatocardium_ciliatum 1 -163 45.8 83.9 -24.3 -118 -26.7 68.1 12.4 -108 46.7
cirripedia 1 -54.9 4.88 -21.6 -0.258 -0.374 3.4 -13.5 17.6 6.25 -11.9
cistenides_granulata 0.19 -2.04 -0.126 -0.328 0.207 0.579 -0.919 -0.194 -0.613 -1.01 0.117
cossura_longocirrata -5.53 -1.89e+15 -2.01e+14 -3.16e+14 -7.08e+13 5.72e+14 -4.38e+14 -1.47e+14 2.18e+14 1.31e+14 -4.14e+14
crassicorophium_bonellii 1 -1804 475 -1567 -50.6 539 127 747 -1319 530 56.4
crenella_decussata -4.3 -2.64e+15 2.6e+14 -2.04e+15 5.86e+14 7.3e+14 2.7e+14 6.91e+14 -1.66e+15 -3.77e+14 5.93e+14
cumacea 1 -71.1 30.5 -31.3 -32.1 -14.7 5.09 18 -7.97 21.5 10.3
cyclocardia_borealis 1 -77.8 -2.56 -29.7 9.63 43 -5.21 13.2 -55.7 24.6 -9.86
cyrtodaria_siliqua 1 -180 54.8 -26.2 -54 -32.2 22.3 -58.4 -71.2 50.8 29.4
diastylis_rathkei 0.18 -345 -1.1 -0.0255 0.391 0.633 -1867 0.199 -1.42 0.443 -0.211
diastylis_sculpta 0.35 -105 -1.55 -0.145 -65.8 0.374 -451 -1.91 -0.532 1.18 -0.335
diastylis_sp 1 -225 91.9 -168 51.8 204 7.32 27.8 -20.2 -18.1 21.9
echinarachnius_parma 0.58 -9.71 0.153 -6.73 -0.342 2.87 -1.22 -6.79 -0.728 3.29 -4.78
edotia_montosa 1 -417 -152 58.9 38.1 -173 -17.5 86.6 45.9 52.7 -218
ennucula_tenuis 0.3 -0.31 1.31 -0.17 0.0216 0.187 0.439 0.109 -0.774 -0.599 1.03
eteone_sp 0.33 -349 1.44 -5.05 0.473 2.26 -1880 1.36 -1.04 1.04 0.173
euchone_sp 0.42 -57.7 -3.29 -3.03 -56.5 2.37 -172 -3.2 0.249 3.15 -1.97
eudorella_emarginata 0.45 -232 0.0201 -0.224 1.69 3.34 -1233 0.579 1.16 -1.12 0.548
eudorellopsis_integra 0.31 0.821 0.583 0.89 -0.533 -0.298 -0.0988 1.04 -0.409 -1.37 2.14
euspira_pallida -5.53 -4.09e+15 -6.24e+13 -5.44e+14 3e+14 7.08e+14 4.18e+14 5.27e+14 -3.86e+14 -8.54e+13 -2.37e+14
glycera_capitata 1 -87.1 24.4 -64.1 21.9 21.3 20.4 -37.3 29.3 46.4 -0.209
glycera_sp 0.35 -6.09 -2.18 0.228 0.972 1.93 -9.26 -0.228 0.231 0.223 -1.73
goniada_maculata 0.43 -321 1.86 -0.0531 -2.03 -0.0395 -1748 -0.939 0.972 -1.04 1.01
guernea_prinassus_nordenskioldi 1 -656 163 -573 22.5 234 -33.2 15.2 -13.5 330 -476
halacaridae_spp 1 -157 -23.2 -123 -11.4 54.9 -74 32.9 -60 -3.26 21.2
haminoea_solitaria 0.33 -106 -0.166 2.83 -61.4 -1.43 -444 2.9 0.0562 -6.72 1.72
hardametopa_carinata 0.42 -57.7 -3.29 -3.03 -56.5 2.37 -172 -3.2 0.249 3.15 -1.97
harmothoe_sp 0.56 -154 2.83 -6.23 3.98 10 -780 0.576 -5.54 6.48 -2.86
harpacticoida 0.13 -0.205 -0.621 -0.295 -0.253 -0.503 -0.53 0.243 0.0159 0.0593 -0.0601
hediste_diversicolor 0.49 -4.76 -5.08 0.523 0.686 2.24 -1.09 0.582 -0.948 -1.05 0.147
heteranomia_squamula 1 -127 -0.595 -25 20.9 41.3 23.5 -129 -37.4 18.8 18.9
hiatella_arctica 1 -191 18 -64.3 -0.669 -16.5 63.7 -184 -8.11 12.1 30
holothuroidea 0.7 -81.9 0.57 -8.04 -122 -1.86 -11.5 -52.3 -7.6 3.6 8.5
idotea_phosphorea 1 -1101 -887 29.4 102 59.1 -87.5 -91.1 -46.7 -3.92 128
ischyroceridae_spp 1 -266 -7.97 177 -26.3 -93.4 45.8 -366 63.6 31.2 52.9
ischyrocerus_anguipes 0.57 -349 -11 3.37 0.306 -1.26 -1819 -4.91 0.496 1.44 -0.727
isopoda 1 -520 183 142 -285 -380 -64 65.7 48.5 -172 70.8
lacuna_vincta 1 -317 -175 -111 52.1 58.2 0.527 72.2 -80.5 -87.3 71
lamprops_fuscatus 0.14 -2.66 -0.185 -0.866 -0.776 0.13 -0.753 -0.712 -0.466 0.723 -0.313
lamprops_quadriplicata 1 -114 6.83 -34.4 -8.6 -54.3 -3.45 56.8 12.1 -38.6 26.1
lepeta_caeca 0.44 -355 -0.0369 -4.1 1.25 3.46 -1911 -1.55 -0.861 0.179 -0.452
leucon_leucon_nasicoides 0.31 -0.484 1.24 1.21 -0.789 -1.17 0.285 0.0632 0.471 -0.938 1.48
littorina_littorea 0.3 -358 -2.67 0.937 0.503 0.586 -1931 -0.126 -0.176 0.122 -1.57
lumbrineridae_spp 1 -255 98.3 -189 40.9 164 21.5 -86 65.4 -10 -43.3
lysianassidae_spp 1 -598 9.21 -225 -94.3 224 81.6 -647 -125 149 -32.4
macoma_calcarea 0.16 1.47 -0.396 0.169 -0.0437 0.871 -1.15 -0.0234 -0.468 -0.691 -0.409
maera_danae 1 -1544 -951 125 -544 -45.2 -10.2 -502 -1.57 162 -59.1
maldane_sarsi 1 -635 263 -321 203 508 43.1 17.1 68.9 323 -200
maldanidae_spp 0.19 -13 0.798 -0.135 -41.7 0.569 0.448 -0.229 -0.183 0.658 0.143
monoculopsis_longicornis 0.42 -40.1 1.14 -4.09 -64 1.33 -55.6 -8.88 -4.09 0.962 2.13
muculus_musculus_discors 1 -157 -23.2 -123 -11.4 54.9 -74 32.9 -60 -3.26 21.2
mytilus_sp 0.27 -4.91 0.0251 0.761 0.132 0.81 -3.22 -6.67 0.42 0.665 -0.27
nematoda 0.38 -1.31 0.679 -2.4 0.0473 0.528 -2.77 1.46 -0.613 -1.1 0.142
nemertea 1 -85.7 23.5 -65.6 19.6 17.9 1.23 -31.3 26.3 36.7 13.3
neoleanira_tetragona 1 -105 -18.4 59.3 -15.4 -57.8 -7.62 32.3 7.49 -75.5 33.1
nephtyidae_spp 0.23 -142 0.674 0.361 -60.9 -1.68 -668 2.34 0.0639 -1.65 0.574
nephtys_caeca 0.3 -365 -1.24 0.598 0.495 -0.449 -1976 0.139 0.754 -0.322 -1.65
nephtys_incisa 0.29 -1.41 1.23 0.331 0.136 0.254 0.728 -0.638 -0.257 -0.426 1.01
nephtys_sp 0.17 -131 1.13 -1.98 -51.2 0.892 -611 0.96 -0.265 0.289 0.0405
nuculana_minuta 0.2 -315 0.959 -1.9 0.603 2.12 -1702 -1.03 -0.124 0.379 -0.0358
nymphonidae_spp 1 -127 -0.595 -25 20.9 41.3 23.5 -129 -37.4 18.8 18.9
oenopota_sp 0.37 -344 -1.05 -1.77 0.544 2.85 -1855 0.946 -2.19 -0.403 -0.212
oligochaeta 0.2 -141 -0.439 -0.0873 -57.6 0.499 -671 -0.447 0.149 0.511 0.23
ophelia_limacina 0.14 -355 0.887 -1.6 0.475 0.516 -1918 -0.497 0.181 1.13 0.258
opheliidae_spp 1 -294 111 -150 41.8 121 54.6 49.9 26.9 9.76 -174
ophiopholis_aculeata 0.42 -70.4 0.461 -3.78 -57.6 3.11 -259 -0.0764 -1.84 -0.87 -0.106
ophiura_robusta 0.58 -271 0.814 -7.89 -0.2 2.89 -1414 -0.624 -4.71 0.573 -1.08
orchomenella_minuta 0.26 -225 -0.299 2.08 -0.302 0.508 -1190 -5.64 -1.49 -0.324 0.992
ostracoda 0.12 -1.17 0.442 -0.387 -0.993 0.207 0.441 -0.651 -0.369 0.281 0.139
pagurus_sp 1 -150 -9.76 4.93 1.68 40.3 -16.3 -135 -3.05 -39.9 20.9
pagurus_pubescens 0.65 -248 2.23 -6.86 0.211 -1.01 -1256 -11.5 0.156 -0.971 2.41
pandalus_montagui 1 -60.1 9.47 -26.5 9.25 28.5 -1.94 5.58 -40.6 31.3 10.1
parathyasira_equalis 0.33 -106 -0.166 2.83 -61.4 -1.43 -444 2.9 0.0562 -6.72 1.72
parvicardium_pinnulatum 0.3 -7.54 1.52 -6.57 0.618 3.82 -1.01 -4.88 0.701 2.35 -0.957
periploma_leanum 0.27 -140 0.697 -2.32 -46.1 1.53 -671 1.3 0.0568 0.432 0.474
philine_lima 1 -150 -9.76 4.93 1.68 40.3 -16.3 -135 -3.05 -39.9 20.9
philomedes_sp 0.6 -22.9 4.82 -2.47 -0.0119 -1.34 -49.2 5.03 -1.89 -2.45 -3.69
pholoe_longa 0.42 -57.7 -3.29 -3.03 -56.5 2.37 -172 -3.2 0.249 3.15 -1.97
pholoe_sp 0.08 -0.00379 -0.17 0.743 -0.0626 0.0978 0.172 -0.46 -0.56 0.0319 -0.103
phoxocephalus_holbolli 0.12 -1.58 -0.503 0.15 -0.103 -0.555 0.0334 -0.629 0.55 -0.192 -0.159
polynoidae_spp 0.11 -348 0.0281 0.397 -0.229 0.466 -1891 -0.542 -0.0922 -0.26 0.126
pontogeneia_inermis -5.53 -3.15e+15 -9.8e+13 -1.97e+15 1.86e+14 1.13e+15 -1.29e+13 -6.04e+14 2.02e+14 9.76e+14 -2.94e+14
pontoporeia_femorata 0.32 -1.46 -0.344 0.0831 0.137 0.764 -1.26 -0.38 0.108 0.535 0.693
praxillella_praetermissa 0.24 -16.5 0.357 -1.05 -36.1 1.92 -18.9 0.881 -0.0499 0.553 -0.499
propebela_turricula 0.54 -124 5.07 -4.46 -23.8 -0.0915 -604 5.07 -2.3 0.682 2.05
protomedeia_fasciata 0.16 -331 0.165 0.655 0.0182 -0.135 -1793 -1.78 0.832 -0.237 0.664
protomedeia_grandimana 0.15 -0.341 0.463 0.668 -0.0449 -0.226 0.126 0.157 -0.511 0.072 0.656
puncturella_noachina 1 -782 203 -670 245 749 -301 -367 -218 256 54.8
quasimelita_formosa 0.12 -1.46 0.552 -0.294 0.106 0.933 -0.908 -0.387 0.0131 0.227 -0.512
quasimelita_quadrispinosa 1 -266 -7.97 177 -26.3 -93.4 45.8 -366 63.6 31.2 52.9
retusa_obtusa 0.31 -18.1 -1.79 -0.334 -50.3 0.532 -1.16 0.408 0.206 0.711 -1.47
sabellidae_spp 0.28 -16.4 -0.679 -1.04 -48.5 0.59 0.292 -0.377 0.936 0.0393 0.0683
scoletoma_fragilis 0.36 -16.1 -1.2 0.588 -45 2.07 0.536 -0.643 -1.27 0.878 -0.873
scoletoma_sp 0.55 -70 -2.64 -6.67 -71.5 2.66 -220 2.17 -2.21 1.17 -2.92
scoloplos_sp 0.29 -83.3 1.77 -0.708 -49.4 -0.917 -337 3.94 0.317 -5.78 1.45
serripes_groenlandicus 0.71 -162 5.74 -4.15 12.4 25.3 -661 -0.398 16 -1.63 -0.966
sipuncula 0.23 -13.5 0.278 0.224 -41.4 0.869 0.559 0.699 -0.138 -0.247 -0.387
solamen_glandula 1 -77.8 -2.56 -29.7 9.63 43 -5.21 13.2 -55.7 24.6 -9.86
solariella_sp 0.5 -213 2.09 -4.74 1.17 4.01 -1103 -5.09 0.0733 -0.624 -3.83
strongylocentrotus_sp 1 -206 1.86 -170 20.3 106 -206 114 -101 -40 33.8
tachyrhynchus_erosus 0.29 -83.3 1.77 -0.708 -49.4 -0.917 -337 3.94 0.317 -5.78 1.45
thracia_septentrionalis 0.19 -1.85 -1.25 -0.279 0.418 0.739 -0.209 -0.101 -0.189 -0.633 -0.337
thyasira_gouldi 0.18 -1.34 0.921 -0.638 -0.00046 0.697 -0.803 0.0839 -0.606 0.0371 0.337
thyasira_sp 0.29 -83.3 1.77 -0.708 -49.4 -0.917 -337 3.94 0.317 -5.78 1.45
trichotropis_bicarinata 0.36 -27.1 3.74 -0.9 -36.2 -2.37 -31.2 -4.88 1.56 1.09 -2.29
turritellopsis_stimpsoni 0.28 -266 1.18 0.308 -0.409 -1.52 -1422 -1.81 -1.18 -0.36 1.45
yoldia_myalis 0.09 -324 0.515 0.4 -0.126 -1.16 -1750 0.68 0.089 -0.565 0.445
Exposure indices
McFadden’s pseudo-R2 and variables coefficients for each taxon
  Pseudo-R2 intercept depth aquaculture dredging runoff sewers structures shipping fisheries
aceroides_aceroides_latipes 0.09 -1.7 -0.101 0.509 -1.02 0.362 0.0924 0.0161 0.617 0.182
akanthophoreus_gracilis 0.23 -1.55 0.566 0.823 -0.0484 0.97 -0.0888 -1.18 0.651 0.429
ameritella_agilis 0.17 -3.08 0.108 -0.0254 0.973 0.343 0.449 -2.15 -0.956 0.056
ameroculodes_edwardsi 0.51 -13.5 2.14 -0.507 5.28 4.02 -6.11 -3.79 -5.65 -1.75
ampelisca_vadorum 1 -302 34.1 -8.99 -26.5 -303 -198 427 68.8 25.1
amphipoda 0.07 -0.617 0.171 0.189 0.666 0.736 0.515 -1.63 -0.155 0.097
anonyx_lilljeborgi 0.22 -3.75 -1.05 0.13 -2.23 -1.39 -1.49 2.13 0.283 -1.67
anthozoa 1 -98.4 11.6 -9.1 -9.56 -104 -33.5 100 8.97 -63.3
arcteobia_anticostiensis 1 -117 -19.1 68.4 5.86 28.7 36.2 -15.2 5.82 -111
arrhoges_occidentalis 1 -130 9.69 6.16 -15.3 -26.5 -97.8 67.8 83.7 -45.5
astarte_sp 0.64 -107 2.33 -262 -8.75 0.667 -6.03 6.58 -6.35 -0.132
axinopsida_orbiculata 0.14 -1.44 0.412 -0.217 0.776 0.247 -0.289 -0.472 -0.283 0.329
axiothella_catenata 1 -3807 153 -17.5 1451 -2304 1318 -3107 -519 -99.3
bathymedon_longimanus 0.61 -13.9 -6.85 4.32 1.2 -0.191 -0.68 4.3 -2.79 -2.65
bathymedon_obtusifrons 1 -1160 -27 304 -1808 137 429 17.2 -72.5 -20.6
bipalponephtys_neotena 0.16 2.08 0.63 0.554 0.144 0.591 1.08 -0.986 1.03 0.127
boreochiton_ruber 0.73 -94.7 -2.45 -5.36 -55.5 -115 -47.3 167 -12 5.77
brachydiastylis_sp 1 -246 30.1 -22.2 -18.1 -82.4 -185 200 47.2 30.6
byblis_gaimardii -3.38 -3.1e+15 -1.39e+14 -4.06e+14 1.2e+15 8.83e+13 -1.66e+15 -6.38e+14 3.23e+14 2.5e+14
cancer_irroratus 1 -1039 -189 708 135 417 1328 -526 -174 362
caprella_septentrionalis 0.55 -37.2 -9.89 -61.9 -4.64 1.67 -2.55 0.383 1.29 -2.6
chaetodermatida 0.22 -2.13 1.16 0.572 -0.0153 2.2 0.603 -1.81 0.0916 0.53
chionoecetes_opilio 1 -194 44 30.9 -45.4 -30.8 -59.5 55.6 55 -224
chlamys_islandica 1 -39.6 -10.9 1.93 2.89 -5.51 6.96 -2.32 2.21 9.57
chone_sp 1 -162 -3.62 71.9 -104 72 113 -28.2 -15.7 41.6
ciliatocardium_ciliatum 1 -192 61.2 -97.1 25.4 -124 -50.8 123 -7.72 -2.98
cirripedia 1 -268 -113 -18.4 -166 -181 -47.7 200 -56.1 -3.88
cistenides_granulata 0.18 -1.87 -0.214 -0.0253 -0.598 -0.918 -1.11 1.59 -0.038 0.342
cossura_longocirrata 1 -349 -193 -58.1 -15.4 -225 -260 238 221 15.4
crassicorophium_bonellii 0.31 -120 -1.06 -317 -1.03 -3.81 -3.57 5.07 2.27 0.0328
crenella_decussata 0.62 -26.7 0.288 -0.67 -23.6 -30 -18.4 46.6 5.13 1.05
cumacea 1 -459 33.9 47.5 30.6 -673 261 236 -110 -35.5
cyclocardia_borealis 1 -98.4 11.6 -9.1 -9.56 -104 -33.5 100 8.97 -63.3
cyrtodaria_siliqua 1 -627 192 -41.7 123 -260 -331 367 127 -59.6
diastylis_rathkei 0.33 -4.08 -1.86 -2.37 0.946 -0.425 -2.83 0.933 0.346 -0.289
diastylis_sculpta 0.47 -5.72 -2.44 -1.48 -2.54 -0.97 -3.83 3.65 1.97 -0.604
diastylis_sp 1 -162 54.5 -43.7 46.1 -47.6 7.91 26.2 -117 -56
echinarachnius_parma 0.26 -2.72 -0.672 0.103 -0.587 -0.575 -2.03 2.45 0.112 0.216
edotia_montosa 1 -1147 -393 196 27.6 -561 177 923 -851 -377
ennucula_tenuis 0.31 -0.482 1.34 0.855 -0.0468 0.79 -0.118 -0.336 0.937 0.317
eteone_sp 0.4 -1364 0.13 -349 -2729 -3.84 0.599 1.6 -0.0926 0.311
euchone_sp 1 -992 -393 328 -948 153 261 -56.6 123 -20.5
eudorella_emarginata 0.48 -42 -0.795 1.06 -0.0731 -14.2 0.826 6.78 0.151 -71.3
eudorellopsis_integra 0.3 0.69 0.52 0.122 0.384 0.0411 -0.118 -0.947 1.83 -0.305
euspira_pallida 1 -182 42.4 -5.37 87.1 88.4 16.8 -269 -1.81 -19.3
glycera_capitata 1 -2662 -49.4 3.15 -2252 -4181 252 3194 12.9 -24.7
glycera_sp 0.57 -82.7 -3.97 -155 -7.83 -3.14 -4.23 4.32 -0.163 -42.5
goniada_maculata 0.27 -0.667 1.24 0.588 -0.457 0.462 0.451 -0.544 0.477 0.277
guernea_prinassus_nordenskioldi 0.26 -85.4 0.106 -220 -3.89 -3.94 0.403 5.31 -2.31 0.047
halacaridae_spp 1 -39.6 -10.9 1.93 2.89 -5.51 6.96 -2.32 2.21 9.57
haminoea_solitaria 1 -931 87.2 165 92.3 -268 318 516 -972 -199
hardametopa_carinata 1 -992 -393 328 -948 153 261 -56.6 123 -20.5
harmothoe_sp 0.54 -1305 0.71 -349 -2594 -11.4 -1.38 11.3 0.632 -0.108
harpacticoida 0.22 -0.277 -0.569 -0.29 -0.963 1.41 -0.331 -0.685 0.429 0.193
hediste_diversicolor 0.38 -77.6 -1.97 -205 0.0973 -1.63 0.445 1.92 -0.952 -1.79
heteranomia_squamula 1 -5266 -135 374 -4481 -5222 -2947 8586 1727 631
hiatella_arctica 0.64 -55 0.371 -115 -9.07 -10.5 -4.12 19.2 -7.02 0.00213
holothuroidea 1 -205 2.99 -23.2 -37.2 54.6 -126 81.2 -108 -10.9
idotea_phosphorea 0.65 -198 -5.95 -14.8 -392 9.08 2.12 -10.3 1.56 1.59
ischyroceridae_spp 1 -448 -19 243 -102 107 181 146 -36.8 -13.8
ischyrocerus_anguipes 0.69 -87.3 -23.3 -118 -29.5 -12.8 -14.6 21.5 -6.61 -22.1
isopoda 1 -67.9 10.5 28.9 15.8 68.4 23.7 -58.7 10.6 20.1
lacuna_vincta 0.52 -33.8 -25.3 -0.578 -7.15 4.31 -4.2 -2.06 6.97 -0.769
lamprops_fuscatus 0.14 -2.44 -0.396 0.913 -0.0614 0.689 -0.179 0.357 0.552 0.426
lamprops_quadriplicata 1 -468 -386 155 72.1 -9.51 85.7 -92.8 64.8 -44.2
lepeta_caeca 0.35 -9.02 -0.534 -1.32 -11.7 -2.1 -1.97 4.51 -2.71 0.302
leucon_leucon_nasicoides 0.27 -0.66 1.06 0.344 -0.344 -1.19 0.486 0.877 0.6 -0.243
littorina_littorea 0.5 -53.4 0.756 -123 -3.65 -0.0384 -0.578 4.51 -7.73 -0.934
lumbrineridae_spp 0.38 -13.8 0.497 -1.85 -10.6 -15 -0.711 18.1 -5.6 -0.346
lysianassidae_spp 1 -266 -45.5 28.2 -26.8 -212 -83 334 -34.4 27.4
macoma_calcarea 0.08 1.56 0.221 0.0465 0.457 1.86 -0.286 -1.62 0.155 0.229
maera_danae 1 -162 -3.62 71.9 -104 72 113 -28.2 -15.7 41.6
maldane_sarsi 1 -235 28.1 77.5 -87.2 -235 49.4 307 68.6 52
maldanidae_spp 0.2 -1.6 0.813 0.624 -0.694 -0.132 0.815 0.281 0.484 -0.187
monoculopsis_longicornis 1 -203 32.3 14.7 121 173 3.02 -240 -68.5 30.7
muculus_musculus_discors 1 -39.6 -10.9 1.93 2.89 -5.51 6.96 -2.32 2.21 9.57
mytilus_sp 0.29 -4.2 0.606 -0.117 -1.58 0.0799 -0.439 2.86 -3.37 -0.471
nematoda 0.21 -0.733 0.36 0.214 -1.48 0.674 -0.384 -0.54 -0.0541 0.0889
nemertea 0.57 -130 -0.875 -94.6 -82.5 -39.9 6.51 -20.8 1.45 -0.395
neoleanira_tetragona 1 -430 -277 168 99.8 -16.8 218 -178 19.8 -7.44
nephtyidae_spp 0.56 -9.44 1.13 0.382 6.83 6.22 -0.0256 -12.1 -3.48 1.36
nephtys_caeca 0.21 -3.41 -2.1 -0.518 0.0188 -0.624 -1.88 1.09 0.156 -0.0857
nephtys_incisa 0.24 -1.41 1.28 0.374 -0.302 -0.026 0.254 -0.0929 0.524 -0.136
nephtys_sp 1 -2901 433 673 201 518 255 -1154 -73.9 -5291
nuculana_minuta 0.3 -7.52 0.678 0.246 -12.3 -3.06 -0.353 4.66 0.859 0.581
nymphonidae_spp 1 -5266 -135 374 -4481 -5222 -2947 8586 1727 631
oenopota_sp 0.19 -114 -0.699 -308 -0.782 -0.0296 -1.58 0.5 0.245 -0.03
oligochaeta 0.2 -93.3 -0.69 -251 -0.218 0.477 0.379 -0.382 0.855 0.0514
ophelia_limacina 0.32 -126 0.824 -322 -14.3 -1.53 -1.95 3.57 2.26 -0.0963
opheliidae_spp 1 -1660 359 -76.1 -745 -1651 -1107 2600 415 16.3
ophiopholis_aculeata 1 -1008 199 244 -328 -873 -506 1408 706 246
ophiura_robusta 0.4 -10.3 -0.29 -0.898 -8.34 -8.41 -5.65 14.4 -0.242 0.631
orchomenella_minuta 0.26 -99.2 0.902 -261 -3.32 1.08 -0.133 0.587 -0.711 -0.558
ostracoda 0.11 -1.1 0.365 0.28 -0.577 1.21 -0.549 -0.648 0.679 0.108
pagurus_sp 1 -117 -19.1 68.4 5.86 28.7 36.2 -15.2 5.82 -111
pagurus_pubescens 0.59 -18.6 0.401 -0.353 -8.36 -12.3 -3.65 22.7 -13.1 -4.95
pandalus_montagui 1 -452 -220 -50.4 105 -72.4 -306 73 200 -222
parathyasira_equalis 1 -931 87.2 165 92.3 -268 318 516 -972 -199
parvicardium_pinnulatum 0.37 -16.4 -0.319 -0.492 -27.6 -4.86 -2.15 9.56 -2.89 -0.571
periploma_leanum -7.18 -2.73e+15 4.16e+14 7e+14 4.99e+14 7.01e+14 -3.6e+13 -1.13e+15 1.17e+15 -5.92e+12
philine_lima 1 -117 -19.1 68.4 5.86 28.7 36.2 -15.2 5.82 -111
philomedes_sp 1 -369 116 -86.4 90.6 -67.9 -206 114 94 38.7
pholoe_longa 1 -992 -393 328 -948 153 261 -56.6 123 -20.5
pholoe_sp 0.07 -0.0314 0.00619 -0.00222 -0.333 -0.198 -0.966 0.805 0.386 -0.462
phoxocephalus_holbolli 0.15 -1.65 -0.474 -0.124 -0.228 0.288 -0.144 0.0247 -0.887 0.0627
polynoidae_spp 0.13 -1.51 0.591 -0.412 -1.22 -0.0976 0.172 -0.00209 -0.239 -0.773
pontogeneia_inermis 1 -299 -60.8 11.3 -208 -0.565 -98.2 72.4 62.6 -281
pontoporeia_femorata 0.23 -101 -0.163 -277 -0.0145 -1.09 0.779 0.695 -0.206 -0.776
praxillella_praetermissa 0.38 -7.15 0.414 3.95 -0.51 -2.9 2.79 3.96 1.43 -3.58
propebela_turricula 0.58 -1438 4.85 -0.575 -3155 0.524 -0.329 -4.52 3.71 0.393
protomedeia_fasciata 0.07 -2.13 -0.204 -0.346 0.635 -0.486 -0.431 0.234 -0.111 0.169
protomedeia_grandimana 0.13 -0.364 0.895 0.165 0.209 0.353 0.932 -0.513 -0.177 -0.295
puncturella_noachina -6.48 -3.47e+15 -4.26e+14 -8.31e+14 2.89e+14 -2.38e+15 -1.63e+15 1.6e+15 2.39e+14 -2.36e+13
quasimelita_formosa 0.19 -1.49 0.946 1.28 -0.177 1.6 1.29 -1.24 -0.00995 0.392
quasimelita_quadrispinosa 1 -448 -19 243 -102 107 181 146 -36.8 -13.8
retusa_obtusa 0.41 -27.1 -2.52 -56.3 0.0073 -0.815 1.87 2.71 0.796 0.941
sabellidae_spp 0.29 -3.91 -1.63 -1.35 -2.24 -2.78 0.00835 2.37 -0.0696 -0.186
scoletoma_fragilis 0.47 -109 -0.793 -288 -1.53 -5.59 -1.4 6.31 -0.971 -5.15
scoletoma_sp 1 -169 -40.9 -14.7 66.8 -36.3 -94.6 34.1 -12.7 15.2
scoloplos_sp 1 -413 148 225 166 436 318 -448 -153 86.7
serripes_groenlandicus 1 -1327 0.0648 55.2 -6.51 -1576 -797 1452 907 -62.1
sipuncula 0.11 -1.97 0.514 0.0693 -0.15 -1.23 0.446 0.963 -0.126 -0.857
solamen_glandula 1 -98.4 11.6 -9.1 -9.56 -104 -33.5 100 8.97 -63.3
solariella_sp 0.57 -17 2.85 0.878 -10.3 -10.7 -11.3 22 5.99 2.85
strongylocentrotus_sp 1 -257 0.014 -64.9 -6.41 -170 -168 269 -31.1 21.9
tachyrhynchus_erosus 1 -413 148 225 166 436 318 -448 -153 86.7
thracia_septentrionalis 0.13 -1.78 -1.41 -0.11 -0.197 -0.313 -0.909 0.375 0.416 0.0511
thyasira_gouldi 0.26 -1.69 1.11 0.576 -1.63 0.834 0.0589 -0.215 0.855 0.287
thyasira_sp 1 -413 148 225 166 436 318 -448 -153 86.7
trichotropis_bicarinata 1 -182 62.5 5.81 60.7 61.2 -31.4 -63.1 -21.1 -57.4
turritellopsis_stimpsoni 0.37 -1413 0.735 -1.13 -3112 0.482 -3.1 3.5 -3.08 -0.318
yoldia_myalis 0.36 -23.8 0.526 0.434 -41.9 -13.3 -3.26 17.9 2.61 -0.913

2.1.2. Density data

We considered density data with a quasiPoisson distribution. Diagnostics are here.

All parameters
Pearson’s pseudo-R2 and variables coefficients for each taxon
  Pseudo-R2 intercept depth om gravel silt clay arsenic cadmium copper mercury aquaculture dredging runoff sewers structures shipping fisheries
aceroides_aceroides_latipes 0.24 -16.2 -0.17 -0.289 -50.2 0.241 -5.35 0.295 -0.671 0.474 0.166 0.696 -1.35 -0.495 0.342 0.385 0.303 0.0639
akanthophoreus_gracilis 0.53 -1.01 0.647 -0.517 -0.69 -0.478 0.215 1.13 -0.909 0.294 0.471 0.803 -1.16 1.07 -0.377 -0.737 0.602 0.503
ameritella_agilis 0.35 -5.78 0.218 0.917 0.111 -0.956 0.881 -1.73 0.921 0.614 -3.28 0.489 0.413 1.17 1.57 -5.19 -0.0779 0.0714
ameroculodes_edwardsi 1 -405 100 -116 12.9 43.5 3.3 -380 23.7 221 -133 6.25 209 83.5 14.6 -330 -159 -87.6
ampelisca_vadorum 1 -56.7 -5.21 17.4 7.13 0.0427 5.09 19.7 -15.1 -37 -3.94 -0.416 -6.89 8.04 18.3 -17.3 12 6.55
amphipoda 0.29 -0.245 0.117 0.512 0.163 0.363 -0.146 -0.1 0.65 -0.835 -0.0236 0.0749 0.25 0.91 -0.182 -1.09 0.197 0.0681
anonyx_lilljeborgi 0.76 -385 -1.75 0.274 1.5 -0.11 -2067 -4.01 -0.745 1.85 -0.785 0.436 -4.95 -3.45 0.459 1.83 0.0664 -4.38
anthozoa 1 -34.1 -3.62 0.00819 0.942 8.33 -1.75 -0.481 -7.06 -0.0811 -5.43 -4.48 -0.76 -8.64 -1.09 4.72 2.06 -1.49
arcteobia_anticostiensis 1 -45.7 -0.999 -17.8 5.41 11.8 -2.07 0.0791 4.88 -0.821 1.13 11.1 1.53 5.52 -5.33 2.92 3.35 -20.7
arrhoges_occidentalis 1 -45.8 -0.64 1.83 -3.43 -15 -1.14 -6.78 3.71 1.03 -0.754 -12.7 3.66 -11.1 -12.5 2.68 10.1 -13.7
astarte_sp 1 -138 31.1 -32.4 -1.94 14.1 3.54 66.2 -9.05 -32.9 -104 -9.25 -26.3 50.5 37.2 -38.9 -31.7 3.15
axinopsida_orbiculata 0.56 -1.24 0.694 -0.0738 -1.02 -0.739 0.133 -1.69 0.87 1.04 0.144 0.026 0.787 -0.385 -0.839 0.594 -0.212 0.397
axiothella_catenata 1 -43.7 6.25 2.63 -16.4 -10.7 -1.16 2.54 2.29 -5.76 18.4 -12.3 -13.3 -11.2 -7.5 21.3 -12.6 1.13
bathymedon_longimanus 1 -219 -90.4 15.6 29.4 17 -345 -71.3 -21.9 20.3 -83.4 77.1 54.5 -61.5 150 -15.7 -12.6 16.7
bathymedon_obtusifrons 1 -180 -26.6 11.8 -10.2 -6.94 -6.78 -0.29 5.48 30.8 -119 56.4 -153 20.5 78.9 4.67 -8.18 -13.1
bipalponephtys_neotena 0.41 2.58 -0.585 0.0555 0.136 0.692 -0.631 -0.253 0.232 -0.271 -0.101 0.357 -0.307 0.454 -0.0537 -0.185 0.184 -0.127
boreochiton_ruber 1 -127 -14.8 -31.9 14.8 31.6 -86.1 17.1 -39.5 3.82 26.9 -12 -9.11 -42.6 -29.1 93.8 -76.5 2.16
brachydiastylis_sp 1 -63.3 8.49 11.2 5.19 4.58 7.63 26 -0.372 -47.5 3.37 -4.79 -13.1 -0.689 -12.8 18.3 28.8 8.75
byblis_gaimardii 1 -382 -103 245 -5.81 -116 47.9 -142 91.4 6.04 11.5 -60 -95.3 -225 -378 351 34.6 45
cancer_irroratus 1 -72.1 1.66 -34.3 20.2 25.4 -1.81 20 -0.0444 -32.1 1.37 18.2 12.2 27.3 44.8 -28.3 3.19 10.9
caprella_septentrionalis 1 -448 -176 -1.01 111 21.6 4.58 -566 -18.8 211 -104 19.1 -19.8 -162 134 -53.5 -40.6 -111
chaetodermatida 0.54 -4.34 1.66 -0.336 -1.2 -0.798 1.33 -6.75 0.329 1.7 -0.386 0.579 0.819 0.722 1.77 -2.79 0.395 0.0996
chionoecetes_opilio 1 -82.5 19.3 2.96 3.56 2.41 13.4 -55.3 -3.6 31.5 3.35 8.24 -9.41 -19.8 -25.7 15.1 13 -54.3
chlamys_islandica 1 -31.7 -6.1 -1.09 -0.29 2.27 -1.68 0.0531 -1.37 0.326 -0.915 -0.921 0.722 -4.11 1.04 0.762 0.99 3.81
chone_sp 1 -81.1 -1.83 14.3 4.25 -3.23 -1.82 23.4 12.1 -18 3.6 28.6 -63.7 47.4 30.1 -7.31 -7.47 21.9
ciliatocardium_ciliatum 1 -43.6 2.24 17.1 -7.25 -24.4 -8.18 -0.521 4.07 -1.51 -0.442 -8.95 -0.672 -20.8 -3.27 12.6 0.116 1.37
cirripedia 1 -30.9 0.498 0.497 -0.11 -1.32 0.124 -1.41 6.12 -6.29 -0.518 0.233 0.0684 -4.27 0.422 6.64 3.48 1.68
cistenides_granulata 0.53 -2.25 0.33 -1.53 -0.012 0.63 -0.485 1.29 -0.493 -1.33 0.36 -0.106 -1.2 -0.658 -0.57 2.28 -0.632 0.065
cossura_longocirrata 1 -59.7 -12.8 -5.31 7.79 17.7 17.4 -13.4 16.9 7.08 0.225 0.795 4.74 17.1 -41.1 -11.4 24.5 -0.566
crassicorophium_bonellii 1 -67.5 10.7 -18.8 -5.7 -3.16 2.81 11 -26.2 60.5 2.64 9.46 -11.7 -10.9 -39.5 10.1 24.5 4.6
crenella_decussata 1 -124 16 -71.7 6.5 27.1 -2.05 -51.7 -64.6 59.8 20.4 6.46 -34.9 -80.1 -15.9 80.9 0.262 -13.3
cumacea 1 -35 9.17 -6.98 -6.29 -3.8 2.18 1.03 -7.59 11.3 1.13 1.95 4.78 0.349 8.16 -12.4 -5.66 -4.37
cyclocardia_borealis 1 -34.5 -3.83 0.168 0.976 8.71 -1.66 -0.479 -7.37 -0.245 -5.63 -4.8 -0.95 -9.11 -1.32 5.15 2.12 -1.53
cyrtodaria_siliqua 1 -57.4 12 -18.8 -0.641 -12.8 1.34 -16.9 -14.4 30.1 4.08 23 20.3 23.8 20.4 -35 1.75 -2.3
diastylis_rathkei 0.82 -261 -1.63 -0.429 -0.132 -0.405 -1369 -3.21 0.575 2.38 0.222 -20.1 -0.039 -4.58 -3.06 4.02 0.322 -0.788
diastylis_sculpta 0.42 -120 -3.96 -2.53 -83.9 1.2 -499 2.32 -0.778 -0.446 0.483 -1.75 -2.54 -1.61 -3.66 4.45 0.894 -1.81
diastylis_sp 1 -63 15.6 -3.01 8.47 20 8.12 -0.459 15.6 0.969 -8.25 -22.8 4.65 -12.9 -13.8 16.9 -12.9 1.22
echinarachnius_parma 0.98 -11.4 0.527 -8.21 -1.09 2.97 -12.8 -3.93 -1.07 2.89 -5.74 2.12 0.206 0.368 2.9 -1.38 0.9 -0.279
edotia_montosa 1 -73.3 0.647 -2.84 -17.6 -20.4 -4.52 -2.41 1.26 19.5 -39.1 -17 -0.327 -30.5 9 15.9 -23.5 -4.02
ennucula_tenuis 0.5 -0.124 0.77 0.242 -0.153 0.0631 0.298 0.654 -0.485 -0.793 0.571 0.388 -0.544 0.568 -0.334 0.0838 0.361 0.381
eteone_sp 1 -134 -2.55 -64.7 -4.05 0.14 -5.34 4.36 -106 29.1 1.63 -21.7 62.7 -21.5 90.4 -112 -10.4 -7.02
euchone_sp 1 -262 -63.3 -26.8 -48.7 6.42 -14.1 96.8 5.73 -30.9 -75 123 -128 110 102 22.3 0.695 -17.2
eudorella_emarginata 1 -676 -384 96.2 154 203 -687 315 -126 -482 67.3 62.1 -282 -253 166 169 71.8 45.1
eudorellopsis_integra 0.6 1.72 -0.0878 0.584 0.111 0.124 0.103 1.12 0.55 -1.59 0.589 -0.0869 -0.315 1.33 -0.371 -0.29 0.8 0.301
euspira_pallida 1 -82.5 8.83 13.6 15.3 27 16.1 -28.1 7.08 -4.3 -42 -2.8 30.2 43.6 11.6 -85.8 23.1 1.26
glycera_capitata 1 -49.6 5.6 -26.3 7.72 4.36 8.18 -31 11.3 29 -4.78 3.57 8.6 -7.27 2.31 -8.07 2.03 -3.96
glycera_sp 1 -483 -174 -12.8 139 27.6 -42.7 -26.3 -36.2 38.5 10.9 -5.38 -466 -137 -53.1 100 17.3 -373
goniada_maculata 0.43 -329 0.805 0.233 -0.697 -0.404 -1789 -1.43 0.26 0.23 0.0601 0.102 0.188 -0.446 0.192 0.0567 0.439 0.0782
guernea_prinassus_nordenskioldi 1 -106 15.5 -67.7 -35.6 20.8 -9.71 31.7 -17.1 10.8 -39 0.152 -29.4 7.97 7.35 18.4 -22.3 -2.01
halacaridae_spp 1 -31.7 -6.1 -1.09 -0.29 2.27 -1.68 0.0531 -1.37 0.326 -0.915 -0.921 0.722 -4.11 1.04 0.762 0.99 3.81
haminoea_solitaria 1 -58 -8.78 38.4 -4.5 -26.2 -6.82 11.6 8.84 -17.6 -6.03 24.3 -14.9 25.1 7.56 -7.01 12.3 7.73
hardametopa_carinata 1 -184 -30.5 18.5 3.38 -8.62 -5.6 -7.24 9.15 27.4 -119 61.5 -154 38.2 73.4 -8.22 1.11 -11.4
harmothoe_sp 1 -198 53.8 -109 73.4 186 -20.5 -65.6 52.9 64.1 -108 -77.2 42.1 -25.6 15.1 -37.8 8.69 -8.9
harpacticoida 0.41 0.0616 -0.577 0.0274 0.349 -0.00405 -0.348 1.18 0.0586 -1.28 -0.134 -0.402 -0.74 0.667 0.0612 0.0301 0.141 -0.535
hediste_diversicolor 0.82 -28.4 -4.19 1.32 0.566 0.889 0.0597 2.03 -0.0708 -4.42 3.53 -48.8 -3.04 -1.77 0.893 7.23 -3.59 -7.36
heteranomia_squamula 1 -44.4 -2.27 -12.9 11.6 13.4 0.0951 12 -0.491 -9.21 -4.57 4.95 -9.83 8.04 -8.28 12.3 6.65 -9.91
hiatella_arctica 1 -58.3 3.2 -15.8 -2.95 -11 4.74 -12.6 -4.4 -2.55 3.62 -3.14 -10 -18.4 7.31 20.4 4.11 -1.62
holothuroidea 1 -51.4 -13.4 14.9 1.86 -5.86 3.13 9.73 -24.9 8.88 -0.855 0.0326 -11.9 19 -6.26 -12.2 2.01 4.7
idotea_phosphorea 1 -145 -30.4 46.1 46 -2.45 -9.92 16 -5.53 0.782 -106 35.7 -12.7 80.6 123 -86.2 -5.4 36.7
ischyroceridae_spp 1 -65.8 -10.2 38.8 4.03 -7.38 3.24 17.7 10.9 -29.9 5.74 18.1 -28 46.3 -12.1 8.27 -0.62 4.54
ischyrocerus_anguipes 1 -215 -79.7 42.7 35.2 -41.3 27.3 -202 11.1 71 -36 -50.6 -110 -90 -31.8 21.5 34.5 -97.8
isopoda 1 -33.5 0.139 12.2 -3.32 -9.66 -2.21 4.22 3.69 -7.37 3.24 2.42 -2.78 11.8 -2.58 -5.4 4.48 5.39
lacuna_vincta 1 -112 -65.5 -39.7 19.3 27.4 -27.5 -67.7 -17.9 22.9 -13.2 7.38 32.2 -48.1 38.4 -10.9 13.8 -6.16
lamprops_fuscatus 0.17 -2.71 0.327 -0.858 -0.352 -0.22 -0.37 -1.71 -0.35 1.54 -0.405 0.954 0.0295 1.28 0.715 -1.38 0.273 0.437
lamprops_quadriplicata 1 -52.7 2.48 1.71 -11.9 -29.3 -9.45 10.1 2.46 -7.03 5.53 11.9 1.54 -2.69 22.6 0.234 1.06 7.44
lepeta_caeca 1 -330 3.6 -102 30.4 72.4 -17.1 -324 -13 109 -62.6 -24.6 -159 -104 51.1 51.2 -108 -17.6
leucon_leucon_nasicoides 0.49 -0.415 1.01 0.497 -0.169 -0.165 0.0935 0.0798 -1.53 0.933 0.346 0.203 -0.338 -1.14 -0.00949 0.726 1.1 0.63
littorina_littorea 1 -137 6.33 60.5 16.9 -23.4 -3.23 46.1 11.9 -41.7 -30.4 -19.4 -71.2 74.3 8.16 16.3 -66.8 4.19
lumbrineridae_spp 1 -68.5 15.2 5.62 7.67 14.4 6.2 -0.823 22.9 -8.2 -8.55 -27.7 2.2 -25.6 -21.8 35.4 -5.89 2.97
lysianassidae_spp 1 -51.7 -16.9 4.13 6.8 9.34 -0.687 -1.59 -18 15 -15.9 17.4 -3.1 2.86 11.9 -1.6 7.22 5.42
macoma_calcarea 0.44 1.17 -0.0535 -0.25 -0.297 0.111 -0.0287 -0.318 -0.32 0.554 -1.1 0.296 0.0792 0.707 0.724 -0.905 -0.158 0.0156
maera_danae 1 -73.9 -4.38 15.6 5.29 -2.4 -1.56 20.9 12.1 -20 9.48 29.9 -52.2 48.8 31.1 -14.2 -1.03 22.1
maldane_sarsi 1 -68.2 15.9 -10.5 10.5 7.78 0.814 7.71 13.8 20.1 -12.1 7.5 -20.4 1.48 12.7 15.7 5.88 14.5
maldanidae_spp 0.43 -13 0.747 0.544 -43.7 -0.267 -0.23 0.374 -0.927 1.02 -0.000389 0.759 -0.855 -0.188 -0.495 0.611 1.64 -0.00221
monoculopsis_longicornis 1 -93.1 5.4 26.1 10.3 10.7 14.3 23 -55.5 6.52 5.94 -0.426 14.1 54.5 24.4 -125 -0.989 3.39
muculus_musculus_discors 1 -31.7 -6.1 -1.09 -0.29 2.27 -1.68 0.0531 -1.37 0.326 -0.915 -0.921 0.722 -4.11 1.04 0.762 0.99 3.81
mytilus_sp 1 -8.54 -1.65 -0.768 -1.9 0.554 -3.5 -7.89 -1.67 0.0967 1.94 -0.866 -0.0648 -5.64 2.23 5.37 -4.66 -0.59
nematoda 0.77 -1.95 0.53 -3 0.241 0.138 -0.674 -5.15 -0.315 3.37 -2.17 0.691 0.237 0.109 1.27 -3.09 0.381 -0.173
nemertea 1 -51.2 7.31 -27.5 6.38 2.4 2.1 -36.3 13.2 28.9 1.61 2.69 13.3 -1.94 0.702 -15.4 2.41 -1.18
neoleanira_tetragona 1 -43.8 3.75 2.08 -16.6 -4.89 0.732 13.1 -2.67 -24.8 2.67 -0.022 -3.05 6.11 13.3 -9.63 0.239 0.436
nephtyidae_spp 0.88 -98.9 3.68 -6.28 -47.7 0.152 -418 5.92 -2.04 1.37 1.81 0.542 4.65 4.56 2.88 -5.69 -9.56 0.702
nephtys_caeca 0.23 -352 -1.56 0.813 0.711 -0.557 -1900 1.92 0.621 -1.16 -0.691 -0.342 -0.223 0.828 -2.2 1.02 -0.32 0.042
nephtys_incisa 0.44 -2.09 1.35 0.473 -0.132 -0.25 0.486 -1.7 -0.844 1.54 0.268 0.836 -0.102 1.03 -0.574 -1.01 1.34 -0.234
nephtys_sp 1 -163 47.7 -68.1 47.4 93.5 33.8 -142 46.8 18.6 -46.9 -11.5 60.2 12.2 11 -118 53.3 -49.8
nuculana_minuta 0.32 -248 0.801 -1.64 0.547 2.13 -1313 -1.4 -0.324 -1.33 -0.133 0.369 -10.1 -2.13 0.501 3.68 1.55 0.377
nymphonidae_spp 1 -43.2 -1.82 -15.1 10.5 15.5 -1.19 12.5 -2.16 -10.7 0.203 3.98 -10 6.96 -7.75 13.2 7.81 -7.4
oenopota_sp 0.7 -374 0.0788 -1.64 0.832 4.14 -1409 10.8 -0.906 -7.4 3.02 -309 -4.98 2.64 -3.77 5.78 -0.333 1.27
oligochaeta 0.28 -258 -1.34 1.65 -70.6 -0.879 -857 2.7 -0.264 -1.77 1.33 -223 -2.44 0.413 0.0683 2.29 -0.914 0.801
ophelia_limacina 0.66 -145 2.95 0.981 2.11 -1.03 -1.8 -10.2 1.63 4.97 -1.4 -360 -10.1 -3.44 -4.52 2.1 6.68 -4.47
opheliidae_spp 1 -97.4 17.9 27 12.7 10.7 10 21.9 16.1 -39.1 -1.04 -45.5 -24 -14.2 -38 62.7 -9.98 13.8
ophiopholis_aculeata 1 -65.2 -9.44 9.6 5.09 13.5 -90.2 7.97 -14.4 -5.99 -16.4 11.9 -6.83 8.2 6.85 -5.57 13.1 9.99
ophiura_robusta 1 -133 23.6 -56.7 -12.7 7.44 19.1 50.5 -93.7 13.4 25.5 -2.92 -45.2 -27.2 -13.6 63.6 -31 4.74
orchomenella_minuta 1 -75.1 -11.9 31.3 -3.51 -23.5 -2.14 27.9 -82.7 -0.999 18.4 -11.3 -37.3 5.98 45 -30.5 -8.85 1.45
ostracoda 0.98 -1.91 0.578 -2.97 -3.53 1.28 -1.74 0.6 -0.751 0.729 -0.574 0.827 -1.52 -0.647 1.55 1.1 -0.47 -0.42
pagurus_sp 1 -45.7 -0.999 -17.8 5.41 11.8 -2.07 0.0791 4.88 -0.821 1.13 11.1 1.53 5.52 -5.33 2.92 3.35 -20.7
pagurus_pubescens 1 -67 4.53 -24.4 8.52 -2.91 10.8 -7.57 -8.93 1.15 25.9 -3.01 -5.52 -12.3 -19.7 33.2 -23.6 -21.2
pandalus_montagui 1 -34.8 1.54 -10.4 3.06 8.4 -1.36 -0.586 -2.17 4.63 5.45 -0.932 4.19 -3.62 -2.83 6.05 -6.46 -0.676
parathyasira_equalis 1 -59.2 -11 38.1 -1.68 -24 -5.78 13.2 7.54 -17.6 -8.82 25.1 -19.6 24.8 9.84 -5.63 10.9 9.18
parvicardium_pinnulatum 1 -341 0.861 -128 28.9 63.8 -47.4 152 21.4 25.5 -193 24.8 -391 70.5 -88.1 143 -37 -14.3
periploma_leanum 0 35.8 -1.4e-14 5.69e-14 -7.33e-14 -1.18e-13 -1e-14 -1.08e-13 2.47e-14 -5.17e-15 3.45e-14 4.56e-15 1.05e-13 -9.33e-14 -8.46e-14 -2.35e-14 5.82e-14 2.61e-14
philine_lima 1 -45.7 -0.999 -17.8 5.41 11.8 -2.07 0.0791 4.88 -0.821 1.13 11.1 1.53 5.52 -5.33 2.92 3.35 -20.7
philomedes_sp 1 -117 45.2 -17.5 5.19 -6.71 12.6 26.7 6.88 -61 2.8 -20.5 41 20.8 8.5 -39.9 2.88 2.09
pholoe_longa 1 -252 -68.5 -22.6 -26.3 9.88 -13.5 69.9 5.14 -24.9 -73.4 129 -101 108 121 4.24 10.7 -12.9
pholoe_sp 0.57 0.0761 -0.65 0.346 -0.00329 0.502 0.385 0.263 -0.528 0.153 -0.269 -0.0895 -0.658 0.096 -1.53 0.847 0.551 0.107
phoxocephalus_holbolli 0.37 -1.33 -0.6 0.0691 0.363 -0.197 -0.257 -0.0274 0.037 -0.214 -0.405 0.271 -0.238 0.45 0.434 -0.375 -0.837 -0.348
polynoidae_spp 0.34 -393 0.00286 -0.238 -0.202 0.185 -2143 0.115 -0.977 0.0312 0.26 -0.213 -1.35 -0.572 0.952 0.355 -0.732 -0.476
pontogeneia_inermis 1 -109 7.1 -34.4 9.55 38.1 -4.2 -67.5 23.9 81.5 -81.6 3.02 -48 -35 -26.1 31 36.7 -3.55
pontoporeia_femorata 0.31 -76.8 0.145 -0.0861 -0.178 1.19 -1.35 0.288 0.409 -0.659 0.711 -212 -0.698 0.108 0.414 1.26 -1.36 0.205
praxillella_praetermissa 0.96 -28.8 1.87 -0.738 -17.2 -1.03 -2.04 -3.7 4.51 1.55 -3.03 10.1 1.01 -12.3 5.18 11 2.77 -33.2
propebela_turricula 0.57 -1437 2.56 -0.439 -22.6 -0.663 -47.8 4.38 -1.13 -1.66 1.56 -0.294 -3132 -1.54 -1.44 3.49 1.71 0.706
protomedeia_fasciata 0.55 -226 0.818 -0.249 1.09 2.44 -1214 -1.35 1.91 -0.0426 0.164 -1.77 0.661 0.302 -3 1.2 0.0754 0.627
protomedeia_grandimana 0.11 0.874 0.461 -0.0105 0.215 0.127 0.0118 0.417 -0.187 -0.799 0.434 0.128 0.0401 1.24 1.05 -1.38 -0.307 -0.0882
puncturella_noachina 1 -71.3 2.16 -28.1 18.3 38.3 -4.02 -90.3 -4.51 59.2 -16.5 -2.94 20.3 -21.2 -4.32 -28.3 12.6 -18.1
quasimelita_formosa 0.53 -2.68 0.787 -0.43 -0.182 1.08 -5.63 -3.05 -0.202 1.55 -1.04 0.728 0.197 0.121 1.08 -1.48 0.628 0.446
quasimelita_quadrispinosa 1 -66.7 -10.5 39.7 4.09 -7.59 3.3 18.1 11.2 -30.6 5.86 18.5 -28.5 47.5 -12.3 8.39 -0.601 4.69
retusa_obtusa 1 -292 -70.6 -60.4 63.4 99.3 37.2 70.1 47.9 -3.76 -48 118 -105 -18.7 -88.9 247 136 -24.1
sabellidae_spp 0.98 -20.2 -1.29 -2.01 -57.3 -2.57 -0.217 -3.25 1.22 2.69 0.496 -2.42 0.602 -4.06 -0.145 2.49 -1.16 -0.382
scoletoma_fragilis 0.93 -64.4 -1.9 -0.0799 -59.8 4.33 -0.0701 -2.12 -1.49 0.00542 1.13 -100 -4.13 -13 -1.57 19 -6.38 -2.3
scoletoma_sp 1 -41.8 -4.09 -14.2 -0.72 6.52 -0.266 -12.9 -1.4 1.04 -3.22 -1.34 17.6 -2.04 4.73 -12.5 3.67 2.66
scoloplos_sp 1 -73.6 24.8 -12.9 -7.67 -20.1 -5.38 9.66 -11.9 21.9 21.2 28.4 13.2 30.1 35.1 -26.4 -31.9 14.7
serripes_groenlandicus 1 -101 -7.95 -1.57 20.2 37.7 -1.63 -92.3 4.55 60.9 -51.7 10.5 35.8 -87.3 26.5 0.26 21.9 -10.1
sipuncula 0.42 -14.8 0.479 0.14 -46.3 0.905 0.331 0.558 0.143 -0.766 -0.1 -0.153 -0.46 -0.158 0.0546 0.588 0.185 -0.466
solamen_glandula 1 -34.1 -3.62 0.00819 0.942 8.33 -1.75 -0.481 -7.06 -0.0811 -5.43 -4.48 -0.76 -8.64 -1.09 4.72 2.06 -1.49
solariella_sp 1 -205 51.7 -48.5 3.97 37.4 15.6 120 -1.92 -164 -2.96 -3.74 -37.1 -2.68 -19.4 94.1 -15.6 30
strongylocentrotus_sp 1 -50.9 -10.9 11.8 8.14 15 11.5 15.9 -8.08 -47.4 2.71 -6 0.322 -0.585 7.49 -5.78 11.1 8.61
tachyrhynchus_erosus 1 -73.6 24.8 -12.9 -7.67 -20.1 -5.38 9.66 -11.9 21.9 21.2 28.4 13.2 30.1 35.1 -26.4 -31.9 14.7
thracia_septentrionalis 0.79 -2.59 -1.52 0.0244 1.05 0.79 1.1 -2.39 -0.215 -0.396 -1.15 0.588 0.104 1.37 0.384 -1.52 1.09 -0.225
thyasira_gouldi 0.58 -2.04 1.08 0.335 -0.283 -0.275 -4.21 1.26 -1.98 0.657 0.489 0.742 -1.22 1.16 -0.0576 -1.28 0.803 0.261
thyasira_sp 1 -92.2 24.9 -27.9 -3.09 6.93 -108 31 -19.6 8.37 13 24.5 -7.06 35.3 22.3 -18.6 -27.4 10.9
trichotropis_bicarinata 1 -62.9 18.9 0.192 1.47 -6.63 5.49 -25.2 18.3 -8.11 -3.89 0.566 25.5 12.8 0.126 -29.7 17.3 -14
turritellopsis_stimpsoni 1 -450 96.1 169 -49.3 -334 15.5 -371 96.5 99.6 109 35.1 109 137 -132 -150 -67.7 -80.9
yoldia_myalis 1 -245 -5.97 17.9 61.2 -36.9 9.42 4.96 14.2 17.8 7.62 60.3 -307 -84.5 20.5 98.2 7.11 -164
Abiotic parameters
Pearson’s pseudo-R2 and variables coefficients for each taxon
  Pseudo-R2 intercept depth om gravel silt clay arsenic cadmium copper mercury
aceroides_aceroides_latipes 0.06 -15.2 0.0384 -0.62 -50.4 0.651 -2.76 0.242 -0.0494 -0.21 0.246
akanthophoreus_gracilis 0.14 -0.416 0.577 -0.769 -1.12 -0.252 0.174 0.729 -0.445 -0.962 0.469
ameritella_agilis 0.19 -3.01 -0.0969 1.23 -0.0701 -1.42 -0.114 0.232 0.635 -0.717 -0.636
ameroculodes_edwardsi 0.12 -293 1.17 -0.791 0.272 0.728 -1576 -2.09 -0.112 -0.487 0.0308
ampelisca_vadorum 1 -78.3 1.16 -2.96 8.05 1.26 11.3 30.9 -29.6 -37.2 21.7
amphipoda 0.17 -0.112 0.133 0.319 0.106 0.336 -0.233 0.0266 0.483 -0.827 0.0287
anonyx_lilljeborgi 0.14 -371 -1.16 0.241 -0.108 -0.0462 -2014 0.391 -0.698 -0.611 -0.0243
anthozoa 1 -51.4 -0.607 -15.2 5.13 22.8 -3.12 6.04 -28.5 13.6 -4.93
arcteobia_anticostiensis 1 -88.6 -3.82 -2.07 -0.364 21.2 -13.9 -66.1 -1.62 -18.7 9.92
arrhoges_occidentalis 1 -137 19.6 -114 -7.74 8.33 12 62.1 -8.85 -45.7 22.3
astarte_sp 0.18 -238 1.86 -3.43 -0.069 1.7 -1257 -1.05 -2.93 0.22 -0.0507
axinopsida_orbiculata 0.18 -0.981 0.793 0.0485 -1.28 -0.694 0.0907 -1.83 0.783 0.909 -0.552
axiothella_catenata 1 -63.8 14.3 8.42 -4.12 -4.23 10.6 -36.9 -12.4 -0.457 18.2
bathymedon_longimanus 0.66 -264 -0.895 0.893 -2.34 0.996 -1401 -7.12 -0.196 -1.43 0.632
bathymedon_obtusifrons 0.99 -32.9 -0.899 -13 -33.3 9.11 -44.3 -3.73 1.94 8.1 -5.21
bipalponephtys_neotena 0.39 2.67 -0.513 -0.136 0.0758 0.793 -0.648 -0.423 0.183 -0.174 -0.0932
boreochiton_ruber 1 -318 -1.3 -10.9 1.31 6.55 -1656 -1.71 -5.47 -2.06 2.06
brachydiastylis_sp 1 -157 55.6 -88.6 8.84 43.5 38.5 33.5 -44.7 -33 16.3
byblis_gaimardii 0.22 -238 1.74 -1.55 -0.186 -0.326 -1258 -0.672 -0.555 -3.92 1.41
cancer_irroratus 1 -112 -36.5 -61.6 17 35.2 -7.94 51.2 -10.3 -49.1 17
caprella_septentrionalis 0.95 -16.1 -12 -1.16 3.1 0.387 -0.98 -3.54 -0.533 0.986 -1.61
chaetodermatida 0.36 -2.79 1.24 -0.697 -1.14 -0.399 0.562 -1.62 0.0373 0.322 0.384
chionoecetes_opilio 1 -2930 1255 -339 -28.9 150 582 -362 -1121 205 41.3
chlamys_islandica 1 -88.5 -11.7 -60.1 -6.72 26.9 -32.2 14.4 -30.1 0.469 10.2
chone_sp 1 -887 -540 69.6 -307 -25.1 -6.87 -283 -0.911 92.4 -33
ciliatocardium_ciliatum 1 -91.2 22.4 39.7 -11.8 -57.2 -13.2 33.1 5.58 -51.3 22.8
cirripedia 1 -40.4 2.59 -10.2 -0.164 -0.386 2.03 -8.19 9.34 2.94 -5.7
cistenides_granulata 0.32 -2.23 0.12 -1.33 0.143 0.673 -0.465 -1.21 -0.353 -0.307 -0.361
cossura_longocirrata 1 -968 -111 -240 323 692 -122 84 251 36.8 -79.4
crassicorophium_bonellii 1 -915 243 -737 -40.1 220 111 475 -718 184 112
crenella_decussata 0.6 -7.32 0.36 -3.68 0.555 1.75 0.851 -0.622 -1.55 -2.31 0.516
cumacea 1 -48.6 15.8 -14.7 -17.2 -8.74 1.61 8.76 -4.29 11.8 5.46
cyclocardia_borealis 1 -52.6 -1.38 -15.2 5.28 23.3 -2.48 6.68 -29.8 12.9 -4.94
cyrtodaria_siliqua 1 -99.8 26.4 -12.9 -28.8 -18.2 9.66 -27.2 -33.8 24.3 15
diastylis_rathkei 0.06 -366 -1.05 -0.574 0.42 0.564 -1986 -0.975 0.341 0.772 -0.0822
diastylis_sculpta 0.11 -123 -0.97 -1.78 -78.4 1.17 -540 -0.688 -0.147 1.21 -1.13
diastylis_sp 1 -126 46.1 -83.2 25.8 103 3.46 9.27 -9.65 -6.36 10.2
echinarachnius_parma 0.57 -4.2 -0.411 -3.28 -0.501 0.801 -1.23 -0.104 -1.2 0.681 -1.74
edotia_montosa 1 -302 27.8 121 -59.5 -244 -72.3 70.4 83.9 5.18 -132
ennucula_tenuis 0.39 0.0813 0.731 0.0837 -0.163 0.0969 0.284 0.335 -0.294 -1.11 0.444
eteone_sp 0.36 -348 1.26 -4.45 0.543 2.11 -1873 1.87 -1.39 0.503 0.291
euchone_sp 0.07 -63.4 -3.17 -2.89 -68.4 2.28 -205 -3.07 0.25 3.02 -1.93
eudorella_emarginata 0.58 -268 0.108 -0.43 1.56 3.12 -1432 0.49 1.13 -0.924 0.413
eudorellopsis_integra 0.49 2.01 -0.0276 0.1 0.0301 -0.0446 -0.0823 0.744 0.355 -0.707 0.557
euspira_pallida 1 -296984 112516 -139870 8073 72089 85747 51192 -256228 33736 107808
glycera_capitata 1 -55.9 11.3 -31.3 11.3 10.4 9.93 -18.2 14.3 22.9 -0.0155
glycera_sp 0.32 -4.33 -2.03 0.0612 0.556 1.1 -3.04 -0.439 0.247 0.434 -0.563
goniada_maculata 0.37 -351 0.815 0.0471 -0.642 -0.249 -1913 -0.554 0.0892 0.132 0.306
guernea_prinassus_nordenskioldi 1 -342 83 -293 13.6 121 -17.5 -7.03 8.71 160 -235
halacaridae_spp 1 -88.5 -11.7 -60.1 -6.72 26.9 -32.2 14.4 -30.1 0.469 10.2
haminoea_solitaria 0.03 -104 -0.156 2.69 -60.5 -1.35 -435 2.66 0.0547 -6.37 1.63
hardametopa_carinata 0.07 -58.8 -3.17 -2.89 -55.7 2.28 -181 -3.07 0.25 3.02 -1.93
harmothoe_sp 0.33 -149 2.44 -5.21 3.31 8.21 -759 0.414 -4.2 5.46 -2.48
harpacticoida 0.28 0.335 -0.97 -0.0308 0.105 -0.0807 -0.374 0.718 -0.00615 -0.722 -0.0916
hediste_diversicolor 0.58 -3.02 -2.58 0.414 0.232 1.12 -0.235 0.554 -0.226 -0.877 0.357
heteranomia_squamula 1 -79.2 -2.19 -10.5 11.7 21.6 12.5 -66.6 -22.3 9.64 10.8
hiatella_arctica 1 -148 11.6 -24.2 -12.4 -42.6 30.8 -107 -11.2 13.3 28.7
holothuroidea 0.82 -67.3 0.826 -8.83 -96.3 1.98 -45.4 -32 -8.03 2.27 5.94
idotea_phosphorea 1 -621 -458 6.92 63.3 8.6 -26.7 -93 -4.17 29 43.1
ischyroceridae_spp 1 -147 -4.11 89.5 -13.3 -46.8 23.4 -187 32 16.1 26.7
ischyrocerus_anguipes 0.45 -356 -8.23 -0.255 -6.42 -0.943 -1888 0.782 -0.266 0.923 -1.86
isopoda 1 -277 93.1 72.5 -146 -194 -32.7 30.6 25 -85.7 36.1
lacuna_vincta 1 -168 -75.2 -76 14.2 37.3 -6.53 51.2 -45.6 -50 33
lamprops_fuscatus 0.06 -2.42 0.149 -1.11 -0.681 0.0652 -0.452 -1.64 -0.218 0.993 -0.0682
lamprops_quadriplicata 1 -70 3.01 -15.5 -4.66 -28.3 -2.37 28.1 6.01 -19.5 13.3
lepeta_caeca 0.28 -362 0.306 -3.31 0.922 2.49 -1949 -1.44 -0.502 0.101 -0.378
leucon_leucon_nasicoides 0.12 0.452 1.22 -0.479 0.261 0.818 0.351 0.139 -1.08 0.893 0.4
littorina_littorea 0.95 -362 -3.62 0.477 0.94 1.64 -1947 0.138 -0.564 0.155 -1.91
lumbrineridae_spp 1 -146 52.2 -99.1 21.3 86.9 11.2 -45.6 34.6 -5.63 -22.3
lysianassidae_spp 1 -315 4.78 -114 -49.2 113 41.2 -327 -62.7 75.1 -17.3
macoma_calcarea 0.32 1.33 -0.253 -0.222 -0.348 0.118 0.0677 -0.395 -0.172 0.422 -0.709
maera_danae 1 -805 -489 62.8 -277 -22.6 -7.05 -256 -0.874 83.6 -29.9
maldane_sarsi 1 -290 111 -152 85 245 16.2 37.4 -74.7 114 -42.3
maldanidae_spp 0.12 -11.9 0.548 -0.344 -43.5 0.612 -0.185 0.124 -0.0376 0.461 0.196
monoculopsis_longicornis 0.16 -39.2 0.922 -3.13 -66.9 0.184 -53.9 -9.41 -2.76 0.493 2.02
muculus_musculus_discors 1 -88.5 -11.7 -60.1 -6.72 26.9 -32.2 14.4 -30.1 0.469 10.2
mytilus_sp 0.99 -9.23 -2.62 1.65 -1.44 0.285 -3.25 -10.6 -0.869 -1.27 1.48
nematoda 0.32 0.186 0.385 -2.43 0.226 0.163 -0.402 0.646 -0.378 -0.768 0.25
nemertea 1 -61.9 13.7 -37.2 11.9 10.4 0.674 -18.6 15.5 22.1 6.77
neoleanira_tetragona 1 -62.6 -1.57 26.7 -8.7 -33.8 -6.55 16.7 6.21 -34.5 15.9
nephtyidae_spp 0.11 -131 0.685 0.0631 -58.6 -1.14 -614 1.34 0.154 -1.41 0.653
nephtys_caeca 0.2 -371 -1.02 0.696 0.601 -0.516 -2009 0.408 0.363 -0.267 -1.59
nephtys_incisa 0.13 -1.08 0.852 0.0221 -0.171 0.0635 0.309 -0.606 -0.79 0.483 0.422
nephtys_sp 0.02 -128 1.06 -1.88 -50.5 0.883 -599 0.902 -0.242 0.237 0.0344
nuculana_minuta 0.06 -335 0.63 -1.24 0.357 1.4 -1814 -0.551 -0.203 0.136 0.0903
nymphonidae_spp 1 -76.7 -0.0205 -12.7 10.2 20.2 11.6 -64.9 -18.8 9.82 8.88
oenopota_sp 0.58 -370 -0.692 -1.48 0.337 1.93 -1997 0.787 -1.69 -0.291 0.000395
oligochaeta 0.06 -113 -1.36 0.775 -58.8 -0.0806 -526 -1.68 -0.00435 0.261 0.193
ophelia_limacina 0.04 -340 0.858 -1.29 0.374 0.612 -1837 -0.327 -0.0552 0.817 0.194
opheliidae_spp 1 -165 58.1 -77.1 21.7 61.9 28.6 27.5 13.6 4.43 -90.5
ophiopholis_aculeata 0.06 -74 0.476 -3.42 -60 2.65 -276 -0.509 -1.37 -0.653 -0.172
ophiura_robusta 1 -197 -0.166 -15.8 0.0379 7.54 -964 2.64 -12.2 2.1 -0.464
orchomenella_minuta 0.04 -249 -0.274 1.88 -0.252 0.533 -1323 -5.29 -1.36 -0.271 0.918
ostracoda 0.96 -1.85 0.983 -5.13 -0.999 3.55 -2.02 -0.0506 -0.394 2.64 -0.948
pagurus_sp 1 -88.6 -3.82 -2.07 -0.364 21.2 -13.9 -66.1 -1.62 -18.7 9.92
pagurus_pubescens 0.82 -304 1.69 -5.05 0.0161 -1.09 -1587 -7.01 0.253 -0.719 1.69
pandalus_montagui 1 -42.9 4.51 -13.3 4.58 14.6 -1.24 2.55 -20.1 15.6 5.17
parathyasira_equalis 0.03 -109 -0.156 2.69 -62.6 -1.35 -466 2.66 0.0547 -6.37 1.63
parvicardium_pinnulatum 0.2 -7.67 1.37 -6.33 0.826 4.38 -0.799 -5.2 0.508 2.02 -1.04
periploma_leanum 0.02 -143 0.372 -1.08 -52.1 0.323 -680 1.35 0.125 -0.223 0.696
philine_lima 1 -88.6 -3.82 -2.07 -0.364 21.2 -13.9 -66.1 -1.62 -18.7 9.92
philomedes_sp 0.67 -57.7 4.36 -2.68 -0.228 -0.709 -245 4.09 -2.05 -2.49 -2.45
pholoe_longa 0.07 -65.6 -3.17 -2.89 -66.6 2.28 -216 -3.07 0.25 3.02 -1.93
pholoe_sp 0.33 0.286 -0.298 -0.0439 -0.0507 0.502 0.111 -0.13 -0.844 0.314 -0.46
phoxocephalus_holbolli 0.23 -1.05 -1.21 0.0802 0.209 -0.129 -0.0841 -0.0371 0.00994 -0.523 -0.112
polynoidae_spp 0.04 -365 -0.163 -0.283 -0.112 0.439 -1991 -0.18 -0.209 0.0605 -0.011
pontogeneia_inermis 1 -412 12 -285 76.8 230 -16.4 -386 34.8 179 -142
pontoporeia_femorata 0.21 0.328 -0.611 0.135 -0.147 1.2 -1.11 -0.403 0.12 0.124 -0.384
praxillella_praetermissa 0.96 -37.7 0.658 -0.983 -33.8 1.53 -140 1.86 -1.05 1.31 -0.663
propebela_turricula 0.36 -126 2.02 -1.72 -45.1 0.589 -595 1.68 -0.819 -0.472 0.96
protomedeia_fasciata 0.09 -330 0.507 -0.398 0.632 1.15 -1794 -1.87 0.791 0.563 -0.39
protomedeia_grandimana 0.01 1.13 0.0112 -0.0648 0.0204 0.0269 0.056 0.151 0.0296 -0.439 0.383
puncturella_noachina 1 -412 103 -342 125 382 -150 -190 -114 133 27.5
quasimelita_formosa 0.16 -1.36 0.989 -1.13 -0.106 1.63 -2.64 -0.567 -0.406 0.594 -0.271
quasimelita_quadrispinosa 1 -150 -4.2 91.8 -13.7 -48.1 23.8 -191 32.9 16.5 27.4
retusa_obtusa 0.4 -18.8 -1.64 -0.605 -53.3 0.249 -1 0.194 0.455 0.558 -1.29
sabellidae_spp 0.88 -17.2 -1.31 -0.773 -53.5 -1.56 -0.176 -1.17 1.32 0.823 0.0281
scoletoma_fragilis 0.31 -15.1 -0.983 0.628 -46.6 1.31 0.209 -0.389 -0.677 0.328 -0.664
scoletoma_sp 0.47 -89.2 -1.95 -4.78 -68.1 1.9 -340 1.7 -1.57 0.876 -2.39
scoloplos_sp 0.03 -91 1.7 -0.666 -52.2 -0.896 -375 3.83 0.313 -5.61 1.41
serripes_groenlandicus 0.54 -168 5.13 -4.02 11.1 23.4 -714 -0.666 14 -0.899 -2.04
sipuncula 0.25 -12.7 0.437 0.00975 -39.2 1.02 0.358 0.253 0.0569 0.0256 -0.197
solamen_glandula 1 -51.4 -0.607 -15.2 5.13 22.8 -3.12 6.04 -28.5 13.6 -4.93
solariella_sp 0.81 -295 1.31 -6.5 0.928 4.38 -1558 -6.52 -1.19 1.19 -1.81
strongylocentrotus_sp 1 -178 -23.5 -123 25.2 83.8 -25.5 27.2 -31.4 31.9 -83.8
tachyrhynchus_erosus 0.03 -91 1.7 -0.666 -52.2 -0.896 -375 3.83 0.313 -5.61 1.41
thracia_septentrionalis 0.61 -2.35 -1.54 0.145 0.77 0.679 1.05 -2.47 -0.408 -0.322 -0.27
thyasira_gouldi 0.24 -1.82 0.87 -0.512 -0.328 0.416 -7.93 0.96 -1.07 -0.419 0.496
thyasira_sp 0.03 -86.3 1.7 -0.666 -48.4 -0.896 -368 3.83 0.313 -5.61 1.41
trichotropis_bicarinata 0.04 -28.2 3.54 -0.802 -39.3 -2.28 -34.7 -4.57 1.47 1.01 -2.13
turritellopsis_stimpsoni 0.08 -286 1.06 0.249 -0.365 -1.33 -1527 -1.63 -1.13 -0.356 1.33
yoldia_myalis 0.03 -340 0.487 0.374 -0.121 -1.09 -1835 0.66 0.08 -0.55 0.423
Exposure indices
Pearson’s pseudo-R2 and variables coefficients for each taxon
  Pseudo-R2 intercept depth aquaculture dredging runoff sewers structures shipping fisheries
aceroides_aceroides_latipes 0.18 -1.41 -0.0305 0.695 -1.5 -0.37 0.297 0.482 0.645 0.101
akanthophoreus_gracilis 0.52 -0.644 0.274 0.656 -0.476 1.28 -0.568 -1.05 0.835 0.516
ameritella_agilis 0.16 -3.03 0.0865 0.0525 0.843 0.527 0.552 -2.14 -0.849 0.0692
ameroculodes_edwardsi 0.38 -12.1 1.97 -0.387 4.91 3.67 -5.68 -3.79 -3.68 -1.49
ampelisca_vadorum 1 -154 12.5 -9.65 -11.2 -130 -90.9 188 23.2 14.1
amphipoda 0.07 0.0138 0.213 0.0684 0.27 0.247 0.163 -0.996 -0.164 -0.157
anonyx_lilljeborgi 0.65 -4.67 -1.33 0.472 -3.41 -1.46 -1.58 2.44 0.169 -3.27
anthozoa 1 -60.3 6.66 -6.27 -4.69 -54.5 -19.4 56.4 4.37 -24.1
arcteobia_anticostiensis 1 -71.2 -9.93 34.6 2.67 13.5 18.5 -6.89 2.98 -54.1
arrhoges_occidentalis 1 -78.4 4.41 2.52 -7.43 -13.3 -49.8 33.8 42.3 -23
astarte_sp 0.39 -119 1.64 -304 -7.12 0.308 -4.2 4.59 -3.89 -0.354
axinopsida_orbiculata 0.17 -0.682 0.38 0.0719 0.412 0.218 -0.451 0.546 -0.617 0.367
axiothella_catenata 1 -2039 81.8 -8.43 772 -1225 703 -1656 -276 -52.7
bathymedon_longimanus 0.75 -9.41 -3.62 2.78 0.0379 -1.28 -0.328 4.45 -2.77 -2.21
bathymedon_obtusifrons 1 -564 -28.4 102 -929 48.2 125 8.28 -11 -25.5
bipalponephtys_neotena 0.13 2.81 -0.487 0.326 -0.385 0.109 0.0582 -0.0889 0.0609 -0.258
boreochiton_ruber 0.21 -19.8 -0.265 -1.21 -13.2 -22.3 -10.8 34.8 -1.24 1.62
brachydiastylis_sp 1 -144 12.6 -11.2 -9.89 -88.1 -91.6 145 22 14.2
byblis_gaimardii 1 -5843 6.93 -15.5 3263 102 -27.1 -7841 -82.4 25.7
cancer_irroratus 1 -537 -95.7 357 68.2 211 671 -267 -88.2 183
caprella_septentrionalis 0.85 -31.8 -17.1 -39 -4.52 2.19 -6.07 -0.627 4.53 -1.21
chaetodermatida 0.2 -2 0.954 0.0177 -0.0804 0.9 0.262 -0.691 -0.164 0.328
chionoecetes_opilio 1 -111 22.7 15.3 -23.4 -15.4 -30.6 28.3 27.6 -111
chlamys_islandica 1 -32.2 -5.24 0.712 1.24 -3.12 3.17 -0.518 0.741 4.78
chone_sp 1 -106 3.18 39.2 -69.1 63.6 49.5 -33.5 -0.978 24.9
ciliatocardium_ciliatum 1 -106 29.6 -48.2 12.7 -59.3 -24 58.2 -5.61 -1.36
cirripedia 1 -153 -57.3 -10.5 -89.5 -97.8 -24.1 110 -32.5 -1.99
cistenides_granulata 0.2 -1.76 -0.0986 -0.289 -1.27 -1.33 -1.51 2.88 -0.51 0.0606
cossura_longocirrata 1 -245 -1.69 60 -36.8 -21.7 -170 121 210 -27.3
crassicorophium_bonellii 0.56 -131 -0.803 -347 -1.1 -2.84 -4.15 4.48 2.95 0.116
crenella_decussata 0.46 -13.8 -0.0364 0.114 -11.1 -13.8 -8.62 22.3 2.28 0.71
cumacea 1 -220 9.96 37.5 14.4 -268 142 79.3 -52.4 -10.8
cyclocardia_borealis 1 -67 9.27 -14.3 -6.26 -67.4 -23.5 70.9 6.49 -16.1
cyrtodaria_siliqua 1 -328 96.6 -7.4 60.9 -125 -167 182 71.5 -28.3
diastylis_rathkei 0.11 -5.19 -1.74 -9.6 -0.455 -2.44 -1.7 2.88 0.448 -0.445
diastylis_sculpta 0.13 -4.36 -2.4 -1.61 -2.76 -0.777 -3.34 2.66 0.615 -0.856
diastylis_sp 1 -91 26.9 -21.2 23.2 -21.6 3.9 10.6 -56.6 -26.8
echinarachnius_parma 0.32 -1.61 -0.813 0.249 -0.913 -1.55 -1.61 2.86 0.331 -0.454
edotia_montosa 1 -703 -171 84.1 12.2 -240 -1.3 483 -523 -277
ennucula_tenuis 0.42 0.0187 0.78 0.439 -0.088 0.584 -0.226 -0.431 0.383 0.315
eteone_sp 0.27 -1400 0.111 -359 -2801 -3.61 -0.0912 2.27 0.213 0.171
euchone_sp 1 -650 -398 329 -168 161 266 -77.3 127 -4.91
eudorella_emarginata 0.53 -65.7 -0.197 6.66 -0.0433 -12.5 -1.64 7.63 1.01 -127
eudorellopsis_integra 0.27 2.01 -0.22 -0.234 0.296 -0.22 0.193 -0.202 0.524 -0.0145
euspira_pallida 1 -99.8 20.5 -2.51 41.7 43.5 7.91 -129 -0.64 -9.43
glycera_capitata 1 -1377 -25.7 0.661 -1152 -2145 130 1636 6.81 -12.8
glycera_sp 0.18 -32.8 -2.6 -2.23 -6.29 -2.25 -2.4 3.25 -0.289 -56.8
goniada_maculata 0.31 -0.5 0.829 0.151 -0.224 0.118 -0.0191 0.153 0.386 0.151
guernea_prinassus_nordenskioldi 0.99 -18.6 0.0799 -9.1 -13.5 -8.44 5.92 13.1 -7 1.31
halacaridae_spp 1 -32.2 -5.24 0.712 1.24 -3.12 3.17 -0.518 0.741 4.78
haminoea_solitaria 1 -460 30.7 83.1 24.7 -146 160 282 -464 -82.5
hardametopa_carinata 1 -515 -205 168 -469 78.7 132 -29.1 64.9 -11.1
harmothoe_sp 0.27 -1349 0.594 -354 -2688 -10.4 -1.01 10.2 0.527 -0.0714
harpacticoida 0.15 0.416 -0.694 -0.496 -0.0803 0.174 -0.0837 -0.123 -0.0271 -0.306
hediste_diversicolor 0.23 -89.2 -1.04 -240 0.426 -1.17 0.34 0.952 -1 -2.26
heteranomia_squamula 0 -44.3 45.4 -3.54e-14 -2.32e-13 8.27e-12 2.89e-13 -8.18e-12 -2.98e-13 6.28e-14
hiatella_arctica 0.41 -70.4 0.548 -168 -5.21 -6.62 -2.39 12.4 -6.28 -0.288
holothuroidea 1 -115 4.75 -9.15 -20.1 19.2 -68.5 49.2 -37.5 -2.39
idotea_phosphorea 0.87 -266 -16.1 -63.4 -493 8.91 -7.5 -8.41 5.34 0.946
ischyroceridae_spp 1 -239 -9.04 125 -49.5 64.7 95.9 61.5 -18.3 -7.62
ischyrocerus_anguipes 0.3 -44.7 -8.83 -55.5 -15.4 -9.09 -1.34 17.5 -12.7 -8.69
isopoda 1 -46.4 5.25 13.8 8.06 33.8 11.4 -29.6 4.64 9.9
lacuna_vincta 0.97 -43 -33.3 0.499 -8.83 6.1 -6.78 -2.53 9.73 -1.51
lamprops_fuscatus 0.2 -2.13 -0.0334 0.789 -0.582 1.53 -0.0346 -0.306 0.381 0.601
lamprops_quadriplicata 1 -241 -174 81.2 40.7 -4.19 66.6 -61.5 24.7 -20.3
lepeta_caeca 0.22 -7.41 -0.286 -0.89 -10.9 -2.34 -1.64 4.15 -1.22 0.0997
leucon_leucon_nasicoides 0.4 -0.11 1.04 -0.0531 -0.0262 -1.64 0.283 1.36 0.877 0.616
littorina_littorea 0.8 -58.6 0.854 -121 -6.06 0.00671 -4.75 9.34 -14.1 -1.14
lumbrineridae_spp 0.16 -11.8 0.342 -1.72 -10.5 -14.6 -0.737 17.7 -5.32 -0.308
lysianassidae_spp 1 -146 -20.4 14.9 -15.6 -108 -40.9 171 -16.8 14.2
macoma_calcarea 0.18 1.47 -0.0694 0.205 -0.0224 0.906 0.0137 -0.344 -0.219 0.0395
maera_danae 1 -94.6 -1.99 36.6 -52.7 37.8 57.3 -15.6 -7.34 21.3
maldane_sarsi 1 -131 16 37 -43 -123 20.1 160 35.4 25.6
maldanidae_spp 0.33 -0.581 0.73 0.547 -0.726 -0.732 -0.0737 1.21 1.41 -0.193
monoculopsis_longicornis 1 -144 20.8 -3.47 78.3 75 -25.3 -125 -50.2 -13.6
muculus_musculus_discors 1 -32.2 -5.24 0.712 1.24 -3.12 3.17 -0.518 0.741 4.78
mytilus_sp 0.98 -16.9 -2.96 -3.23 -9.38 -9.61 -4.45 20.3 -14.8 -0.253
nematoda 0.18 0.7 -0.555 -0.198 -2.12 -0.814 -1.43 0.83 0.49 0.168
nemertea 0.81 -93.7 -1.68 -50.4 15.8 -68.9 9.7 -17.8 2.2 -0.689
neoleanira_tetragona 1 -230 -139 86 49.9 -6.85 112 -91.2 9.73 -3.92
nephtyidae_spp 0.55 -5.68 0.67 -0.551 3.23 0.745 0.205 -3.35 -3.39 0.479
nephtys_caeca 0.04 -2.84 -1.22 -0.654 0.173 -0.754 -1.58 1.16 0.04 0.0466
nephtys_incisa 0.33 -1.34 1.09 0.56 -0.255 1.27 -0.293 -0.653 0.876 -0.203
nephtys_sp 1 -1509 226 346 98.7 266 127 -588 -30.3 -2714
nuculana_minuta 0.16 -5.24 0.498 0.227 -7.16 -2.35 -0.34 3.62 0.83 0.408
nymphonidae_spp 0.99 -371 -8.93 25.6 -309 -370 -202 605 117 44.1
oenopota_sp 0.36 -120 -0.816 -324 -0.774 -0.347 -1.81 0.754 0.197 -0.181
oligochaeta 0.06 -88.4 -0.653 -244 -0.766 -0.331 0.334 0.924 -0.442 0.269
ophelia_limacina 0.24 -132 0.952 -341 -13.5 -1.48 -1.91 3.13 1.87 -0.579
opheliidae_spp 1 -853 182 -38.3 -377 -838 -560 1317 211 8.41
ophiopholis_aculeata 1 -517 100 121 -162 -435 -253 702 351 123
ophiura_robusta 0.7 -17.9 -0.745 -2.96 -12.2 -20.4 -11.3 32.3 -3.59 1.22
orchomenella_minuta 0.1 -104 0.815 -275 -3.07 1.02 -0.112 0.508 -0.645 -0.507
ostracoda 0.82 -2.18 -0.153 1.92 -4.62 -1.65 3.65 2.49 -1.06 0.206
pagurus_sp 1 -71.2 -9.93 34.6 2.67 13.5 18.5 -6.89 2.98 -54.1
pagurus_pubescens 0.66 -16.3 0.6 -0.294 -6.72 -10.3 -3.01 18.9 -11.3 -4.68
pandalus_montagui 1 -241 -112 -25.5 52.7 -38.6 -154 39 101 -111
parathyasira_equalis 1 -471 30.5 85.6 24.3 -151 165 292 -476 -83.3
parvicardium_pinnulatum 0.12 -14.3 -0.472 -0.101 -23.3 -4.72 -1.8 8.74 -2.07 -0.0718
periploma_leanum 0.32 -996 0.385 403 -3.03 -0.166 5.41 -5.23 648 271
philine_lima 1 -71.2 -9.93 34.6 2.67 13.5 18.5 -6.89 2.98 -54.1
philomedes_sp 0.81 -122 2.69 -312 2.71 -1.86 -4.13 2.16 -0.608 0.416
pholoe_longa 1 -611 -373 302 -178 138 245 -62.6 117 -5.34
pholoe_sp 0.27 0.223 -0.429 -0.199 -0.392 -0.121 -1.39 0.877 0.358 -0.0892
phoxocephalus_holbolli 0.24 -1.1 -0.653 0.181 -0.203 0.428 0.0366 -0.208 -0.908 -0.199
polynoidae_spp 0.12 -0.901 0.194 -0.32 -1.73 -0.439 0.203 0.746 -0.347 -0.476
pontogeneia_inermis 1 -176 -52.1 3.47 -78.8 4.96 -64.3 36 40.9 -152
pontoporeia_femorata 0.11 -88 -0.0333 -246 -0.202 -0.884 0.86 0.92 -1.12 -0.188
praxillella_praetermissa 0.96 -6.49 1.15 3.99 0.869 0.145 3.54 0.173 1.32 -2.72
propebela_turricula 0.53 -1445 2.94 -0.195 -3177 -0.00192 -0.391 -2.33 2.53 0.405
protomedeia_fasciata 0.09 -1.29 0.322 -0.599 0.579 -0.847 -0.751 0.925 -0.502 0.0667
protomedeia_grandimana 0.07 0.941 0.43 0.217 0.223 1.18 0.945 -1.62 -0.203 -0.0666
puncturella_noachina 1 -651 -80.9 -0.718 107 -835 20.8 224 66.6 -10.1
quasimelita_formosa 0.25 -1.2 1.02 0.769 -0.757 0.525 0.725 -0.268 0.477 0.787
quasimelita_quadrispinosa 1 -244 -9.82 128 -51.3 61.4 99.5 67.9 -20 -7.74
retusa_obtusa 0.28 -60 -2.25 -151 -0.0872 -1.2 0.586 2.47 0.463 0.633
sabellidae_spp 0.58 -6.11 -6.11 -1.79 -2.74 -2.19 -1.42 2.26 0.143 0.139
scoletoma_fragilis 0.27 -103 -0.411 -279 -0.537 -3.77 -0.682 3.94 -0.905 -2.21
scoletoma_sp 1 -93.2 -19 -7.2 32.7 -17.2 -45.2 15.8 -5.92 7.42
scoloplos_sp 1 -212 71.7 109 80.2 213 152 -218 -72.2 42.6
serripes_groenlandicus 1 -725 7.55 30.8 -104 -776 -391 702 448 -33.4
sipuncula 0.25 -1.7 0.533 -0.0612 -0.106 -1.28 0.532 0.832 -0.133 -0.906
solamen_glandula 1 -60.3 6.66 -6.27 -4.69 -54.5 -19.4 56.4 4.37 -24.1
solariella_sp 0.26 -10.4 1.28 0.255 -6.71 -8.11 -6.42 15.5 2.74 1.74
strongylocentrotus_sp 1 -142 -0.671 -34.4 -2.92 -86.4 -84.5 135 -16.6 10.9
tachyrhynchus_erosus 1 -212 71.7 109 80.2 213 152 -218 -72.2 42.6
thracia_septentrionalis 0.13 -1.15 -1.06 0.088 -0.336 0.613 -0.57 0.2 -0.358 -0.201
thyasira_gouldi 0.4 -0.993 1.12 0.591 -0.912 1.52 -0.0381 -1.41 0.885 0.232
thyasira_sp 1 -368 82.8 111 -18.6 -289 109 341 -35.2 39.8
trichotropis_bicarinata 1 -101 30 1.38 30.8 30.4 -15.1 -32 -13.6 -28.6
turritellopsis_stimpsoni 0.18 -1453 0.64 -0.979 -3203 0.386 -2.74 3.1 -2.64 -0.266
yoldia_myalis 0.16 -21.8 0.482 0.397 -37.6 -12.1 -2.95 16.3 2.33 -0.842

2.2. Hierarchical Models of Species Communities

This section uses methodology and tools from Ovaskainen et al., with the help of Guillaume Blanchet.

HMSC_PA <- hmsc(data, param, prior, family = "probit", niter = 100000, nburn = 1000, thin = 100)
HMSC_density <- hmsc(data, param, prior, family = "overPoisson", niter = 100000, nburn = 1000, thin = 100)

2.2.1. Presence/absence data

We considered presence/absence data with a probit distribution. Diagnostics are here.

All variables
Model outcomes

Mean of the predictor coefficients estimated by the HMSC model:

Tjur’s pseudo-R2 and variables coefficients for each taxon
  Pseudo.R2 Intercept depth om gravel silt clay arsenic cadmium copper mercury aquaculture dredging runoff sewers structures shipping fisheries
aceroides_aceroides_latipes 0.08 -1.09 -0.00929 -0.0611 -0.202 0.14 -0.204 -0.182 -0.0901 0.0209 0.0478 0.147 -0.336 0.031 0.0968 0.0103 0.225 0.0306
akanthophoreus_gracilis 0.27 -0.987 0.391 0.0233 -0.147 -0.0116 0.257 0.00266 -0.244 -0.113 0.0243 0.309 -0.256 0.172 -0.0167 -0.307 0.346 0.179
ameritella_agilis 0.15 -1.74 0.0241 0.108 -0.0494 -0.256 0.248 -0.0426 0.0986 0.0686 -0.117 -0.00148 -0.165 0.0738 -0.183 -0.206 -0.197 0.0387
ameroculodes_edwardsi 0.13 -2.25 0.421 -0.176 0.0901 0.0811 -0.0985 -0.233 -0.183 0.012 -0.00144 0.0989 -0.179 0.0517 -0.208 -0.0239 -0.0992 -0.0373
ampelisca_vadorum 0.16 -3.55 -0.0494 -0.621 0.265 0.26 -0.303 -0.286 -0.39 0.313 -0.301 -0.202 -0.388 -0.472 -0.109 0.368 -0.196 -0.0266
amphipoda 0.08 -0.467 0.108 0.0771 0.0146 -0.0252 -0.0973 -0.0382 0.184 -0.212 0.283 0.0389 0.0597 0.278 8.87e-05 -0.346 -0.0106 0.0743
anonyx_lilljeborgi 0.07 -1.9 -0.318 -0.181 0.0665 0.177 -0.298 -0.236 -0.248 0.0894 -0.137 0.00251 -0.401 -0.316 -0.0304 0.272 -0.0332 -0.216
anthozoa 0.13 -3.37 -0.187 -0.507 0.102 0.435 -0.342 -0.288 -0.412 0.267 -0.337 -0.234 -0.391 -0.52 -0.159 0.46 -0.231 -0.146
arcteobia_anticostiensis 0.05 -3.15 -0.0431 -0.506 -0.00685 0.278 -0.288 -0.31 -0.189 0.219 -0.182 0.135 -0.291 -0.349 -0.0968 0.365 -0.158 -0.0707
arrhoges_occidentalis 0.04 -3.25 0.0165 -0.564 -0.00971 0.167 -0.272 -0.295 -0.302 0.275 -0.265 -0.168 -0.342 -0.363 -0.155 0.256 -0.0991 -0.105
astarte_sp 0.11 -2.63 0.229 -0.355 -0.0637 0.0747 -0.185 -0.207 -0.394 0.0851 -0.217 -0.185 -0.391 0.0182 -0.145 0.0144 -0.101 0.0133
axinopsida_orbiculata 0.19 -0.928 0.214 0.145 -0.181 -0.117 0.185 0.0489 0.263 -0.252 -0.189 -0.028 0.191 0.225 -0.249 -0.137 0.122 0.267
axiothella_catenata 0.14 -2.64 0.274 -0.28 -0.0438 0.185 -0.167 -0.298 -0.136 0.0547 0.366 -0.0934 -0.29 -0.249 -0.11 0.187 0.0196 0.0012
bathymedon_longimanus 0.11 -2.45 -0.264 -0.291 -0.0636 0.178 -0.278 -0.31 -0.0212 0.0962 -0.281 0.236 -0.0319 -0.16 -0.178 0.442 -0.134 -0.0471
bathymedon_obtusifrons 0.06 -2.9 -0.255 -0.517 0.0421 0.383 -0.396 -0.266 -0.238 0.366 -0.242 -0.117 -0.463 -0.341 0.145 0.474 -0.21 -0.15
bipalponephtys_neotena 0.32 1.43 0.141 0.844 -0.0762 0.144 0.148 0.00701 0.19 -0.691 0.485 0.119 -0.0695 0.473 0.0525 -0.174 0.608 0.163
boreochiton_ruber 0.18 -3.09 -0.294 -0.634 0.121 0.354 -0.381 -0.331 -0.489 0.167 -0.408 -0.122 -0.331 -0.458 -0.142 0.442 -0.3 0.0668
brachydiastylis_sp 0.15 -3.18 0.136 -0.538 0.138 0.16 -0.212 -0.258 -0.358 0.219 -0.247 -0.2 -0.339 -0.328 -0.233 0.193 -0.0262 0.0336
byblis_gaimardii 0.19 -2.8 0.216 -0.363 0.0195 -0.00398 -0.162 -0.232 -0.274 0.108 -0.134 -0.0911 -0.265 -0.0914 -0.309 0.0192 -0.0745 0.187
cancer_irroratus 0.03 -3.22 -0.283 -0.575 0.0669 0.232 -0.405 -0.267 -0.198 0.258 -0.238 -0.176 -0.336 -0.276 0.0523 0.289 -0.378 -0.118
caprella_septentrionalis 0.19 -2.56 -0.556 -0.391 0.0104 0.0708 -0.23 -0.249 -0.14 0.161 -0.212 -0.236 -0.385 -0.0586 0.0988 0.207 -0.514 -0.213
chaetodermatida 0.23 -1.33 0.581 -0.0839 -0.271 -0.25 0.237 -0.131 -0.122 -0.0692 0.0503 0.163 -0.201 0.387 0.114 -0.295 0.149 0.142
chionoecetes_opilio 0.08 -2.82 0.278 -0.323 -0.0313 0.268 -0.182 -0.263 -0.372 0.178 -0.209 0.053 -0.385 -0.251 -0.0799 0.199 -0.00561 -0.0972
chlamys_islandica 0.27 -3.55 -0.157 -0.646 0.0122 0.311 -0.351 -0.296 -0.298 0.343 -0.301 -0.189 -0.294 -0.432 -0.0798 0.356 -0.254 0.191
chone_sp 0.02 -3.13 -0.271 -0.45 0.0139 0.274 -0.351 -0.27 -0.177 0.324 -0.183 -0.167 -0.354 -0.24 -0.0767 0.446 -0.289 -0.0359
ciliatocardium_ciliatum 0.03 -3.19 0.193 -0.483 0.043 0.0686 -0.222 -0.252 -0.275 0.294 -0.154 -0.143 -0.329 -0.341 -0.0818 0.208 -0.155 0.000694
cirripedia 0.13 -3.13 -0.306 -0.439 0.0218 0.27 -0.326 -0.289 0.178 0.076 -0.232 -0.188 -0.215 -0.378 -0.161 0.555 -0.198 -0.0608
cistenides_granulata 0.18 -1.22 -0.0437 -0.0768 0.00538 0.141 -0.199 -0.168 -0.266 -0.225 -0.0819 0.0676 -0.203 -0.118 -0.0654 0.159 0.0306 0.142
cossura_longocirrata 0.01 -2.68 -0.105 -0.196 0.021 0.305 -0.21 -0.144 -0.0436 0.191 -0.0798 -0.136 -0.306 -0.296 -0.148 0.273 0.15 -0.0764
crassicorophium_bonellii 0.03 -2.83 -0.0337 -0.553 -0.0254 0.0425 -0.27 -0.0519 -0.258 0.328 -0.245 -0.153 -0.312 -0.299 -0.116 0.277 -0.0933 -0.0225
crenella_decussata 0.4 -2.92 -0.038 -0.768 0.322 0.17 -0.32 -0.336 -0.673 0.187 -0.464 0.0741 -0.462 -0.48 -0.000125 0.302 -0.205 -0.112
cumacea 0.33 -2.63 0.434 -0.205 -0.0993 -0.0792 0.323 -0.0952 -0.132 0.251 -0.0212 -0.0194 -0.284 -0.175 0.029 0.0877 0.0569 -0.0603
cyclocardia_borealis 0.13 -3.43 -0.185 -0.517 0.0907 0.43 -0.344 -0.285 -0.401 0.274 -0.342 -0.243 -0.389 -0.511 -0.193 0.456 -0.233 -0.15
cyrtodaria_siliqua 0.03 -3.04 0.071 -0.468 -0.0275 0.182 -0.257 -0.255 -0.275 0.235 -0.233 -0.0649 -0.102 -0.159 -0.0527 0.392 -0.154 -0.0595
diastylis_rathkei 0.09 -1.82 -0.405 -0.0318 0.127 0.218 -0.245 -0.148 -0.139 -0.0162 0.0113 -0.308 0.103 -0.0958 -0.438 0.376 -0.109 -0.0573
diastylis_sculpta 0.14 -2.2 -0.488 -0.12 -0.032 0.31 -0.3 -0.262 -0.078 0.204 -0.134 -0.192 -0.355 -0.17 -0.286 0.66 -0.0442 -0.0971
diastylis_sp 0.03 -2.91 0.272 -0.267 -0.00357 0.28 -0.193 -0.236 -0.188 0.152 -0.1 -0.0671 -0.313 -0.239 -0.111 0.256 -0.109 -0.027
echinarachnius_parma 0.3 -2 -0.152 -0.631 -0.131 0.143 -0.34 -0.32 -0.435 0.147 -0.528 0.173 -0.151 -0.0869 0.0293 0.346 -0.0852 0.0395
edotia_montosa 0.13 -3 -0.501 -0.497 -0.0131 -0.0308 -0.409 -0.142 -0.201 0.418 -0.398 -0.205 -0.268 -0.297 -0.0244 0.423 -0.437 -0.196
ennucula_tenuis 0.39 -0.248 0.673 0.197 -0.00627 -0.00788 0.238 -0.0186 -0.238 -0.398 0.268 0.356 -0.0954 0.38 0.0163 -0.193 0.497 0.157
eteone_sp 0.15 -2.42 0.211 -0.543 0.0434 0.139 -0.251 -0.21 -0.391 0.263 -0.218 -0.0984 -0.402 -0.406 0.3 -0.0413 0.142 0.122
euchone_sp 0.02 -3.11 -0.26 -0.439 0.0469 0.302 -0.353 -0.295 -0.184 0.347 -0.208 -0.145 -0.39 -0.273 -0.0577 0.452 -0.211 -0.121
eudorella_emarginata 0.23 -2.13 -0.0857 0.145 0.268 0.373 -0.175 -0.11 -0.0876 0.0402 0.178 -0.15 -0.351 -0.322 -0.0214 0.254 0.278 -0.136
eudorellopsis_integra 0.45 0.358 0.293 0.665 -0.0918 0.0468 0.168 0.179 0.113 -0.649 0.5 0.00051 -0.0909 0.298 -0.258 -0.152 0.772 0.0494
euspira_pallida 0.02 -2.97 0.125 -0.404 -0.0289 0.177 -0.218 -0.217 -0.286 0.22 -0.17 -0.114 -0.34 -0.0566 -0.149 0.115 -0.0684 -0.0612
glycera_capitata 0.09 -2.97 -0.14 -0.553 0.267 0.315 0.0439 -0.239 -0.291 0.295 -0.249 -0.201 -0.39 -0.475 -0.0213 0.35 -0.117 -0.0731
glycera_sp 0.16 -2.33 -0.547 -0.0414 0.0807 0.371 -0.34 -0.201 0.0471 0.172 -0.136 -0.34 -0.293 -0.314 -0.25 0.324 -0.346 -0.218
goniada_maculata 0.4 -0.45 0.652 0.169 -0.36 0.0326 -0.117 -0.25 0.149 -0.39 0.234 0.24 -0.156 0.179 0.123 -0.201 0.37 0.202
guernea_prinassus_nordenskioldi 0.05 -2.97 -0.159 -0.648 0.00397 0.388 -0.411 -0.251 -0.322 0.31 -0.311 -0.15 -0.412 -0.421 0.126 0.449 -0.231 -0.0994
halacaridae_spp 0.28 -3.48 -0.151 -0.631 -0.0105 0.308 -0.352 -0.286 -0.293 0.337 -0.305 -0.176 -0.295 -0.427 -0.0848 0.371 -0.225 0.195
haminoea_solitaria 0.03 -3.11 -0.0218 -0.436 -0.0377 0.164 -0.268 -0.285 -0.215 0.246 -0.202 0.149 -0.297 -0.305 -0.0863 0.3 -0.19 -0.0574
hardametopa_carinata 0.02 -3.1 -0.268 -0.439 0.0472 0.312 -0.353 -0.304 -0.187 0.332 -0.197 -0.142 -0.385 -0.26 -0.0441 0.456 -0.218 -0.123
harmothoe_sp 0.11 -2.68 0.169 -0.326 0.2 0.359 -0.226 -0.23 -0.402 0.163 -0.226 -0.191 -0.418 -0.418 0.0103 0.199 0.0245 0.0322
harpacticoida 0.25 -0.235 -0.291 0.0838 -0.0746 -0.178 -0.221 0.196 -0.0205 -0.282 -0.0896 -0.135 -0.415 0.472 0.0896 -0.226 0.146 0.0402
hediste_diversicolor 0.34 -1.67 -0.766 0.126 0.109 0.495 -0.225 0.0229 0.1 -0.25 0.152 -0.302 -0.123 -0.096 -0.132 0.634 -0.338 -0.172
heteranomia_squamula 0.07 -3.4 -0.189 -0.608 0.132 0.272 -0.373 -0.306 -0.369 0.239 -0.321 -0.13 -0.368 -0.35 -0.00815 0.525 -0.218 -0.11
hiatella_arctica 0.19 -3.04 -0.0455 -0.613 0.0931 0.11 -0.357 -0.321 -0.443 0.145 -0.312 -0.126 -0.47 -0.245 0.113 0.475 -0.203 -0.147
holothuroidea 0.11 -2.72 -0.0655 -0.271 -0.109 -0.0105 -0.229 -0.213 -0.342 0.232 -0.244 -0.15 -0.295 0.155 -0.251 0.102 -0.167 0.0635
idotea_phosphorea 0.14 -2.93 -0.477 -0.492 0.107 0.284 -0.452 -0.236 -0.207 0.166 -0.242 -0.253 -0.352 -0.15 0.109 0.189 -0.536 -0.205
ischyroceridae_spp 0.02 -2.91 -0.172 -0.266 0.0305 0.291 -0.283 -0.258 -0.178 0.252 -0.128 -0.127 -0.39 -0.188 -0.101 0.491 -0.136 -0.12
ischyrocerus_anguipes 0.16 -2.55 -0.646 -0.134 -0.0107 0.107 -0.377 -0.253 0.0971 0.258 -0.201 -0.285 -0.295 -0.2 -0.16 0.356 -0.467 -0.192
isopoda 0.06 -3.1 0.0448 -0.386 -0.0675 0.0773 -0.228 -0.206 -0.213 0.201 -0.151 -0.15 -0.264 -0.0557 -0.21 0.133 -0.157 0.153
lacuna_vincta 0.09 -3.1 -0.411 -0.565 0.0777 0.211 -0.427 -0.27 -0.284 0.181 -0.284 -0.191 -0.393 -0.273 0.0703 0.367 -0.361 -0.138
lamprops_fuscatus 0.09 -1.54 -0.0817 -0.271 -0.149 0.0923 -0.234 -0.203 -0.149 0.0658 -0.169 0.215 -0.0768 0.023 0.0699 0.21 0.109 0.0552
lamprops_quadriplicata 0.09 -3.06 -0.359 -0.575 0.00983 -0.0273 -0.404 -0.0945 -0.181 0.3 -0.264 -0.163 -0.317 -0.3 0.121 0.289 -0.385 -0.178
lepeta_caeca 0.18 -2.61 -0.15 -0.57 0.224 0.325 -0.374 -0.266 -0.44 0.138 -0.334 -0.163 -0.394 -0.334 0.00952 0.205 -0.341 0.0336
leucon_leucon_nasicoides 0.37 -0.342 0.545 0.526 -0.176 -0.141 0.201 -0.00547 0.0701 -0.37 0.376 0.211 -0.216 0.11 0.0597 -0.014 0.448 0.0529
littorina_littorea 0.11 -2.44 -0.384 -0.097 0.00703 0.18 -0.356 -0.17 -0.0961 0.207 -0.177 -0.223 -0.368 -0.0397 0.0243 0.197 -0.474 -0.181
lumbrineridae_spp 0.07 -2.55 0.0551 -0.205 -0.0332 0.258 -0.204 -0.255 0.217 -0.0877 -0.146 -0.131 -0.212 -0.249 -0.155 0.387 -0.0919 0.000961
lysianassidae_spp 0.08 -3.06 -0.218 -0.5 -0.0212 0.28 -0.292 -0.309 -0.353 0.33 -0.401 0.135 -0.283 -0.273 -0.112 0.493 -0.199 -0.0373
macoma_calcarea 0.18 0.83 -0.0325 0.339 -0.125 0.096 -0.206 0.0567 -0.0113 -0.593 -0.0469 0.0381 0.0256 0.566 0.053 -0.342 0.159 0.166
maera_danae 0.02 -3.11 -0.276 -0.443 0.0167 0.278 -0.337 -0.262 -0.163 0.297 -0.187 -0.152 -0.346 -0.225 -0.0702 0.428 -0.289 -0.0226
maldane_sarsi 0.06 -2.48 0.191 -0.0975 -0.00999 0.337 -0.203 0.0255 -0.176 0.182 -0.118 -0.063 -0.36 -0.236 0.113 0.326 0.0121 0.0592
maldanidae_spp 0.24 -0.991 0.387 0.241 -0.199 0.159 0.202 -0.0431 -0.123 0.00586 0.0336 0.227 -0.363 0.0893 0.146 0.0307 0.392 -0.00481
monoculopsis_longicornis 0.14 -2.68 0.159 -0.297 -0.107 0.00973 -0.17 -0.165 -0.305 0.12 -0.0683 -0.154 -0.324 0.25 -0.26 -0.0954 -0.108 0.0737
muculus_musculus_discors 0.26 -3.46 -0.168 -0.593 -0.00786 0.296 -0.336 -0.273 -0.269 0.318 -0.293 -0.185 -0.288 -0.409 -0.115 0.365 -0.235 0.187
mytilus_sp 0.11 -1.57 0.0386 -0.0472 -0.057 0.181 -0.254 -0.302 -0.0648 -0.0619 -0.106 0.0596 -0.22 0.152 0.0368 0.314 -0.274 -0.0689
nematoda 0.49 -0.678 0.313 -0.437 -0.0565 -0.211 -0.285 0.0302 -0.582 -0.109 -0.26 0.264 -0.55 0.0961 0.504 -0.579 0.0369 -0.0475
nemertea 0.13 -2.72 0.078 -0.4 0.218 0.181 -0.22 -0.286 -0.195 0.113 0.263 -0.172 -0.346 -0.339 -0.0675 0.208 0.0892 -0.036
neoleanira_tetragona 0.03 -3.14 -0.241 -0.484 -0.0272 0.04 -0.352 -0.241 -0.152 0.18 -0.201 -0.171 -0.318 -0.215 0.0457 0.202 -0.338 -0.125
nephtyidae_spp 0.2 -1.72 0.14 -0.0346 -0.25 -0.0915 -0.147 0.0767 0.147 -0.0791 0.084 0.011 0.205 0.235 -0.242 -0.204 -0.182 0.293
nephtys_caeca 0.12 -1.84 -0.447 -0.0886 0.0862 -0.0071 -0.264 -0.0989 0.206 -0.0686 -0.21 -0.143 -0.0831 -0.091 -0.282 0.349 -0.141 -0.0275
nephtys_incisa 0.32 -0.886 0.651 0.292 0.0417 0.00537 0.274 -0.167 -0.18 -0.21 0.326 0.18 -0.283 0.139 0.0216 -0.115 0.383 0.00445
nephtys_sp 0.03 -2.55 0.246 -0.321 -0.0353 0.199 -0.178 -0.223 -0.239 0.186 -0.115 -0.0719 -0.383 -0.236 0.0579 0.0918 0.177 -0.109
nuculana_minuta 0.14 -1.8 0.386 -0.265 0.0347 0.268 -0.192 -0.246 -0.333 0.0217 -0.0773 0.102 -0.462 -0.267 0.28 0.103 0.175 0.119
nymphonidae_spp 0.07 -3.38 -0.208 -0.59 0.136 0.283 -0.369 -0.306 -0.354 0.231 -0.291 -0.15 -0.362 -0.342 -0.0514 0.545 -0.218 -0.122
oenopota_sp 0.12 -2.01 -0.2 -0.203 0.0537 0.438 -0.288 -0.111 -0.36 -0.0933 -0.15 -0.238 -0.356 -0.0693 -0.0759 0.188 0.0169 0.0355
oligochaeta 0.14 -1.58 -0.215 0.117 -0.0922 0.253 -0.214 -0.216 0.0677 0.0773 0.0431 -0.104 -0.239 -0.0381 0.024 0.178 0.286 -0.0415
ophelia_limacina 0.07 -2.1 0.237 -0.235 0.129 0.116 -0.161 -0.215 -0.234 0.129 -0.00852 -0.0852 -0.443 -0.163 0.082 0.0619 0.318 -0.0843
opheliidae_spp 0.09 -2.9 0.259 -0.348 0.181 0.253 -0.208 -0.233 -0.285 0.135 -0.166 -0.125 -0.329 -0.323 -0.125 0.233 -0.0844 0.00642
ophiopholis_aculeata 0.15 -3.02 0.0614 -0.473 -0.0378 0.337 -0.262 -0.25 -0.379 0.206 -0.302 0.0691 -0.315 -0.339 -0.0381 0.245 -0.0837 0.181
ophiura_robusta 0.19 -2.95 -0.0488 -0.691 0.00505 0.206 -0.371 -0.313 -0.611 0.164 -0.439 -0.0958 -0.426 -0.412 0.0519 0.359 -0.214 0.028
orchomenella_minuta 0.05 -2.37 0.0597 -0.139 -0.00335 0.201 -0.24 -0.221 -0.397 0.0682 -0.131 -0.107 -0.477 -0.0482 0.103 0.198 -0.0637 -0.123
ostracoda 0.15 -0.723 0.211 0.101 -0.256 0.00773 0.197 -0.17 -0.177 -0.0562 -0.0122 0.108 -0.267 0.317 -0.106 -0.204 0.336 0.0685
pagurus_sp 0.05 -3.14 -0.0553 -0.501 0.00902 0.297 -0.292 -0.325 -0.194 0.221 -0.17 0.129 -0.301 -0.353 -0.0927 0.363 -0.178 -0.0722
pagurus_pubescens 0.12 -3 0.0797 -0.642 0.131 0.0627 -0.306 -0.329 -0.392 0.207 -0.268 0.0434 -0.417 -0.266 0.0741 0.365 -0.264 -0.157
pandalus_montagui 0.1 -2.86 -0.124 -0.263 0.0462 0.385 -0.234 -0.224 -0.0932 0.189 0.233 -0.175 -0.0251 -0.33 -0.217 0.482 -0.0902 -0.0643
parathyasira_equalis 0.03 -3.12 -0.00344 -0.458 -0.0351 0.153 -0.281 -0.271 -0.224 0.256 -0.224 0.154 -0.308 -0.29 -0.0693 0.277 -0.183 -0.0521
parvicardium_pinnulatum 0.05 -2.47 -0.0403 -0.538 0.000133 0.256 -0.288 -0.279 -0.222 0.209 -0.219 0.0428 -0.391 -0.248 -0.0152 0.349 -0.217 -0.113
periploma_leanum 0.1 -2.41 0.127 -0.262 -0.00205 0.258 -0.209 -0.143 -0.181 0.176 0.0205 -0.025 -0.403 -0.275 0.175 0.113 0.332 -0.0891
philine_lima 0.06 -3.12 -0.0714 -0.493 0.0033 0.285 -0.278 -0.319 -0.192 0.214 -0.177 0.147 -0.299 -0.342 -0.0864 0.362 -0.168 -0.0678
philomedes_sp 0.19 -2.65 0.382 -0.402 0.0824 0.0138 -0.12 -0.228 -0.275 0.119 -0.148 -0.178 -0.171 -0.143 -0.241 0.047 0.0827 0.0632
pholoe_longa 0.02 -3.07 -0.267 -0.424 0.0405 0.305 -0.341 -0.282 -0.187 0.326 -0.202 -0.155 -0.388 -0.27 -0.0603 0.457 -0.209 -0.123
pholoe_sp 0.12 -0.0717 -0.0541 0.446 -0.0295 0.038 0.118 -0.0526 -0.143 -0.245 0.0435 0.0788 -0.201 0.254 -0.281 0.0745 0.245 -0.0693
phoxocephalus_holbolli 0.14 -1.04 -0.239 0.0427 -0.0805 -0.119 -0.0741 -0.0724 0.149 -0.125 0.00889 -0.0454 -0.13 0.176 -0.0675 -0.00917 -0.289 0.0311
polynoidae_spp 0.15 -0.942 0.111 0.249 -0.0791 0.156 -0.212 -0.203 -0.213 -0.147 0.0647 -0.114 -0.443 0.0246 0.154 -0.0792 0.0409 -0.176
pontogeneia_inermis 0.03 -3.12 -0.24 -0.434 0.0581 0.329 -0.336 -0.325 -0.189 0.388 -0.175 -0.153 -0.383 -0.268 -0.0818 0.497 -0.208 -0.137
pontoporeia_femorata 0.36 -0.945 -0.163 0.247 0.0571 0.312 -0.258 -0.0196 0.154 -0.117 0.425 -0.187 -0.151 0.0217 0.0861 0.315 -0.0248 -0.143
praxillella_praetermissa 0.12 -1.74 0.0748 -0.0609 -0.063 0.39 -0.198 0.0177 0.0538 0.0237 -0.0463 0.206 -0.0763 -0.162 0.096 0.355 0.253 -0.0389
propebela_turricula 0.17 -2.32 0.599 -0.284 -0.121 0.145 -0.115 -0.192 -0.233 0.124 0.0771 -0.0494 -0.353 -0.19 0.101 -0.0393 0.215 0.1
protomedeia_fasciata 0.15 -1.45 -0.028 0.174 0.0283 0.133 -0.137 -0.195 0.352 -0.239 0.327 -0.169 0.135 0.0524 -0.429 0.23 -0.0249 0.171
protomedeia_grandimana 0.21 -0.267 0.335 0.426 0.0272 -0.016 0.109 0.163 -0.103 -0.311 0.33 0.0737 -0.167 0.404 0.116 -0.0903 0.0934 -0.0452
puncturella_noachina 0.16 -3.29 -0.136 -0.551 0.274 0.363 -0.325 -0.311 -0.448 0.259 -0.311 -0.281 -0.394 -0.561 -0.19 0.382 -0.179 -0.102
quasimelita_formosa 0.16 -0.858 0.326 0.0547 -0.0467 0.185 -0.17 -0.0579 -0.111 -0.115 -0.108 0.336 -0.204 0.257 0.249 -0.111 0.12 0.082
quasimelita_quadrispinosa 0.02 -2.92 -0.174 -0.269 0.022 0.282 -0.293 -0.274 -0.18 0.283 -0.128 -0.129 -0.381 -0.185 -0.0894 0.481 -0.143 -0.11
retusa_obtusa 0.22 -2.21 -0.476 -0.0795 -0.0374 0.318 -0.268 0.0726 0.315 0.0195 -0.135 -0.142 -0.063 -0.0617 -0.282 0.745 -0.0478 0.0308
sabellidae_spp 0.13 -1.77 -0.405 -0.196 -0.0438 0.293 -0.0948 -0.163 0.223 -0.0263 -0.0364 -0.218 -0.296 -0.337 -0.0213 0.407 -0.00791 -0.0841
scoletoma_fragilis 0.21 -1.9 -0.483 0.141 0.0669 0.613 -0.0655 -0.126 -0.184 0.0469 -0.0676 -0.265 -0.346 -0.442 -0.155 0.764 -0.0866 -0.189
scoletoma_sp 0.18 -2.96 -0.23 -0.532 -0.0514 0.279 -0.325 -0.234 -0.153 0.176 -0.258 -0.179 0.101 -0.25 -0.184 0.33 -0.204 0.204
scoloplos_sp 0.03 -3.1 0.117 -0.461 -0.0456 0.173 -0.242 -0.272 -0.219 0.245 -0.193 0.134 -0.291 -0.297 -0.0582 0.291 -0.159 0.00396
serripes_groenlandicus 0.15 -2.51 -0.061 -0.0257 0.298 0.273 -0.174 -0.15 -0.073 0.137 0.113 -0.134 -0.24 -0.295 -0.121 0.309 0.199 -0.12
sipuncula 0.18 -1.2 0.134 0.257 -0.108 0.261 0.211 0.219 -0.0239 -0.157 -0.0526 0.0456 -0.235 0.0499 -0.106 0.21 0.19 -0.0427
solamen_glandula 0.14 -3.38 -0.179 -0.499 0.0875 0.436 -0.349 -0.292 -0.417 0.251 -0.324 -0.237 -0.397 -0.51 -0.171 0.456 -0.209 -0.145
solariella_sp 0.21 -2.55 0.228 -0.414 0.0979 0.293 -0.214 -0.248 -0.44 0.00434 -0.316 -0.0232 -0.34 -0.238 -0.0931 0.213 0.0464 0.201
strongylocentrotus_sp 0.3 -3.31 -0.198 -0.622 0.193 0.405 -0.355 -0.279 -0.48 0.226 -0.405 -0.301 -0.356 -0.548 -0.178 0.354 -0.213 0.135
tachyrhynchus_erosus 0.03 -3.07 0.127 -0.457 -0.0451 0.186 -0.246 -0.262 -0.229 0.25 -0.179 0.126 -0.296 -0.262 -0.0657 0.254 -0.167 0.0139
thracia_septentrionalis 0.14 -1.14 -0.491 -0.121 0.0845 0.231 -0.164 -0.181 -0.0656 -0.225 -0.202 -0.0715 -0.166 -0.157 0.0266 0.136 0.115 -0.00542
thyasira_gouldi 0.27 -0.942 0.525 0.028 -0.0598 0.112 -0.11 -0.111 -0.415 -0.11 0.0138 0.225 -0.49 0.128 0.263 -0.209 0.402 0.0646
thyasira_sp 0.03 -3.09 0.12 -0.462 -0.0394 0.192 -0.244 -0.255 -0.23 0.255 -0.196 0.135 -0.306 -0.274 -0.0567 0.265 -0.172 0.0171
trichotropis_bicarinata 0.01 -2.81 0.181 -0.339 -0.0328 0.174 -0.189 -0.238 -0.162 0.158 -0.127 -0.0941 -0.151 -0.0925 -0.137 0.218 -0.0451 -0.0657
turritellopsis_stimpsoni 0.05 -2.61 0.238 -0.321 -0.0556 -0.00525 -0.176 -0.242 -0.302 0.164 -0.0217 -0.0577 -0.312 -0.0531 -0.154 0.012 -0.176 -0.00852
yoldia_myalis 0.02 -2.26 0.125 -0.199 0.0786 0.116 -0.204 -0.169 -0.215 0.108 -0.0339 0.079 -0.416 -0.231 0.0727 0.235 0.0629 -0.128
AVERAGED 0.14 -2.28 -0.0203 -0.252 0.008 0.183 -0.204 -0.188 -0.183 0.0911 -0.115 -0.0586 -0.28 -0.151 -0.0503 0.213 -0.0564 -0.0201
Ancillary information

Predictive power plot:

95 % confidence interval of the predictor coefficients estimated by the HMSC model:

Variance partitioning:

Trace plots:

Density plots:

Abiotic parameters
Model outcomes

Mean of the predictor coefficients estimated by the HMSC model:

Tjur’s pseudo-R2 and variables coefficients for each taxon
  Pseudo.R2 Intercept depth om gravel silt clay arsenic cadmium copper mercury
aceroides_aceroides_latipes 0.04 -1.03 -0.00225 -0.183 -0.213 0.231 -0.268 -0.174 -0.0186 0.0408 0.126
akanthophoreus_gracilis 0.15 -0.867 0.444 -0.168 -0.0978 0.119 0.221 0.0516 -0.15 -0.309 0.179
ameritella_agilis 0.1 -1.49 0.0224 0.0676 -0.00533 -0.146 0.213 -0.0866 0.164 -0.0891 -0.15
ameroculodes_edwardsi 0.11 -2.1 0.502 -0.41 0.132 0.271 -0.191 -0.152 -0.167 -0.218 0.0277
ampelisca_vadorum 0.14 -3.24 -0.0065 -0.815 0.313 0.425 -0.407 -0.149 -0.274 0.0571 -0.332
amphipoda 0.06 -0.44 0.125 -0.0201 0.0294 0.0342 -0.109 -0.0679 0.159 -0.181 0.284
anonyx_lilljeborgi 0.04 -1.73 -0.318 -0.233 0.0531 0.258 -0.355 -0.185 -0.15 0.0163 -0.16
anthozoa 0.07 -3 -0.108 -0.653 0.093 0.58 -0.402 -0.176 -0.324 0.0414 -0.355
arcteobia_anticostiensis 0.02 -2.85 -0.062 -0.624 0.0646 0.446 -0.379 -0.22 -0.142 -0.0246 -0.191
arrhoges_occidentalis 0.03 -3 0.0553 -0.737 0.0167 0.347 -0.34 -0.157 -0.22 0.0111 -0.274
astarte_sp 0.08 -2.46 0.293 -0.644 -0.017 0.289 -0.271 -0.129 -0.337 -0.0931 -0.186
axinopsida_orbiculata 0.12 -0.887 0.267 0.0224 -0.152 -0.0726 0.154 0.0295 0.117 -0.166 -0.237
axiothella_catenata 0.17 -2.44 0.354 -0.489 -0.0327 0.267 -0.215 -0.192 -0.108 -0.0782 0.377
bathymedon_longimanus 0.05 -2.22 -0.286 -0.319 -0.0446 0.275 -0.351 -0.324 -0.0958 0.0174 -0.336
bathymedon_obtusifrons 0.03 -2.58 -0.302 -0.557 0.0402 0.459 -0.425 -0.222 -0.12 0.274 -0.268
bipalponephtys_neotena 0.24 1.22 0.153 0.744 -0.152 0.0268 0.114 -0.164 0.048 -0.242 0.516
boreochiton_ruber 0.15 -2.73 -0.231 -0.8 0.186 0.518 -0.477 -0.187 -0.432 -0.0397 -0.405
brachydiastylis_sp 0.13 -2.96 0.228 -0.84 0.222 0.366 -0.338 -0.111 -0.317 -0.0509 -0.241
byblis_gaimardii 0.09 -2.44 0.32 -0.635 0.0915 0.193 -0.262 -0.118 -0.268 -0.156 -0.0984
cancer_irroratus 0.02 -3 -0.309 -0.691 0.122 0.405 -0.432 -0.193 -0.132 0.0942 -0.31
caprella_septentrionalis 0.12 -2.2 -0.692 -0.359 0.00899 0.172 -0.184 -0.288 -0.0191 0.102 -0.302
chaetodermatida 0.2 -1.25 0.563 -0.219 -0.254 -0.164 0.234 -0.136 -0.0718 -0.113 0.149
chionoecetes_opilio 0.06 -2.62 0.312 -0.518 0.0151 0.445 -0.277 -0.127 -0.274 -0.0865 -0.169
chlamys_islandica 0.04 -3.06 -0.192 -0.767 0.0747 0.471 -0.434 -0.193 -0.244 0.136 -0.334
chone_sp 0.02 -2.83 -0.348 -0.521 0.0451 0.383 -0.405 -0.274 -0.124 0.229 -0.268
ciliatocardium_ciliatum 0.02 -2.83 0.268 -0.638 0.0681 0.207 -0.267 -0.126 -0.172 0.0206 -0.114
cirripedia 0.12 -2.94 -0.282 -0.557 0.0349 0.31 -0.39 -0.247 0.154 0.0317 -0.338
cistenides_granulata 0.14 -1.16 0.0302 -0.234 0.0548 0.219 -0.258 -0.144 -0.269 -0.281 -0.0338
cossura_longocirrata 0 -2.5 -0.127 -0.341 0.0337 0.411 -0.342 -0.0896 7.34e-05 0.145 -0.123
crassicorophium_bonellii 0.03 -2.59 0.0241 -0.716 0.012 0.164 -0.333 0.0291 -0.18 0.15 -0.218
crenella_decussata 0.35 -2.59 0.0428 -0.908 0.39 0.374 -0.372 -0.119 -0.505 -0.205 -0.361
cumacea 0.33 -2.37 0.43 -0.316 -0.0981 0.0232 0.327 -0.00285 -0.0282 0.0615 0.0226
cyclocardia_borealis 0.07 -3.02 -0.0918 -0.65 0.0978 0.579 -0.404 -0.166 -0.319 0.0411 -0.35
cyrtodaria_siliqua 0.02 -2.91 0.0489 -0.632 0.0061 0.284 -0.328 -0.193 -0.283 0.0998 -0.276
diastylis_rathkei 0.04 -1.62 -0.358 -0.0889 0.108 0.254 -0.343 -0.174 -0.283 0.115 -0.13
diastylis_sculpta 0.09 -1.99 -0.52 -0.192 -0.0615 0.316 -0.415 -0.374 -0.115 0.344 -0.279
diastylis_sp 0.03 -2.66 0.309 -0.455 0.019 0.4 -0.247 -0.148 -0.118 -0.0101 -0.103
echinarachnius_parma 0.27 -1.88 -0.215 -0.68 -0.0946 0.232 -0.407 -0.298 -0.406 0.0708 -0.511
edotia_montosa 0.1 -2.6 -0.559 -0.459 -0.0128 0.0586 -0.413 -0.109 -0.0977 0.304 -0.443
ennucula_tenuis 0.33 -0.205 0.74 0.0226 -0.0275 0.00968 0.185 0.00228 -0.233 -0.333 0.441
eteone_sp 0.05 -2.06 0.239 -0.67 0.0967 0.257 -0.291 0.000678 -0.184 0.0416 -0.0395
euchone_sp 0.02 -2.83 -0.316 -0.507 0.0444 0.408 -0.405 -0.266 -0.121 0.248 -0.289
eudorella_emarginata 0.14 -1.85 -0.0517 0.0353 0.218 0.386 -0.25 -0.0773 0.0189 0.0346 0.24
eudorellopsis_integra 0.31 0.272 0.393 0.429 -0.168 -0.0175 0.084 0.168 -0.0135 -0.368 0.609
euspira_pallida 0.02 -2.86 0.152 -0.667 0.0133 0.387 -0.32 -0.157 -0.226 0.0332 -0.196
glycera_capitata 0.08 -2.68 -0.0756 -0.691 0.288 0.416 0.0447 -0.105 -0.184 0.112 -0.229
glycera_sp 0.09 -2.03 -0.548 -0.058 0.0452 0.441 -0.387 -0.221 0.0777 0.155 -0.288
goniada_maculata 0.36 -0.405 0.741 -0.0255 -0.372 0.0761 -0.153 -0.188 0.169 -0.369 0.419
guernea_prinassus_nordenskioldi 0.03 -2.68 -0.124 -0.73 0.0307 0.475 -0.404 -0.15 -0.215 0.176 -0.298
halacaridae_spp 0.04 -3.05 -0.192 -0.765 0.0686 0.474 -0.424 -0.186 -0.252 0.131 -0.33
haminoea_solitaria 0 -2.7 -0.0439 -0.496 0.0146 0.326 -0.329 -0.178 -0.148 -0.0082 -0.195
hardametopa_carinata 0.02 -2.85 -0.331 -0.524 0.058 0.426 -0.426 -0.273 -0.12 0.244 -0.281
harmothoe_sp 0.07 -2.39 0.23 -0.509 0.248 0.488 -0.282 -0.0945 -0.283 -0.0283 -0.166
harpacticoida 0.13 -0.215 -0.355 -0.0809 -0.118 -0.127 -0.244 0.0126 0.00928 -0.0358 -0.0599
hediste_diversicolor 0.23 -1.44 -0.816 0.161 0.06 0.405 -0.216 -0.124 -0.0123 0.00942 -0.0764
heteranomia_squamula 0.07 -3.23 -0.201 -0.776 0.178 0.419 -0.429 -0.23 -0.309 0.0935 -0.366
hiatella_arctica 0.11 -2.8 -0.0247 -0.751 0.1 0.173 -0.355 -0.252 -0.306 -0.00766 -0.283
holothuroidea 0.05 -2.46 -0.127 -0.429 -0.0599 0.168 -0.336 -0.261 -0.335 0.143 -0.289
idotea_phosphorea 0.09 -2.61 -0.602 -0.526 0.122 0.442 -0.501 -0.232 -0.0939 0.0649 -0.334
ischyroceridae_spp 0 -2.62 -0.225 -0.335 0.0284 0.347 -0.344 -0.282 -0.131 0.216 -0.191
ischyrocerus_anguipes 0.13 -2.18 -0.686 -0.0647 -0.0429 0.146 -0.384 -0.319 0.155 0.249 -0.321
isopoda 0.01 -2.82 0.0435 -0.581 -0.00568 0.242 -0.316 -0.189 -0.195 0.0291 -0.215
lacuna_vincta 0.08 -2.89 -0.461 -0.652 0.0988 0.336 -0.475 -0.227 -0.194 0.063 -0.375
lamprops_fuscatus 0.06 -1.5 -0.154 -0.307 -0.138 0.134 -0.305 -0.235 -0.178 0.128 -0.181
lamprops_quadriplicata 0.06 -2.7 -0.389 -0.603 0.000142 0.0824 -0.385 -0.0292 -0.059 0.139 -0.261
lepeta_caeca 0.12 -2.27 -0.0726 -0.711 0.279 0.5 -0.418 -0.158 -0.327 -0.0639 -0.284
leucon_leucon_nasicoides 0.32 -0.289 0.615 0.366 -0.23 -0.169 0.178 -0.00518 0.114 -0.279 0.56
littorina_littorea 0.06 -2.12 -0.527 -0.0643 0.00236 0.281 -0.38 -0.275 0.00168 0.194 -0.309
lumbrineridae_spp 0.06 -2.38 0.122 -0.365 -0.0246 0.295 -0.248 -0.207 0.192 -0.124 -0.187
lysianassidae_spp 0.05 -2.73 -0.289 -0.539 0.0298 0.422 -0.368 -0.264 -0.296 0.153 -0.437
macoma_calcarea 0.14 0.764 -0.0536 0.227 -0.139 0.115 -0.251 -0.0706 -0.122 -0.311 -0.021
maera_danae 0.02 -2.84 -0.336 -0.511 0.0413 0.384 -0.416 -0.267 -0.104 0.219 -0.276
maldane_sarsi 0.03 -2.24 0.192 -0.22 0.00171 0.406 -0.232 0.0358 -0.0906 0.127 -0.109
maldanidae_spp 0.14 -0.879 0.363 0.103 -0.242 0.169 0.18 -0.0505 -0.0106 0.0543 0.174
monoculopsis_longicornis 0.04 -2.45 0.138 -0.555 -0.0713 0.23 -0.3 -0.188 -0.306 -0.00845 -0.135
muculus_musculus_discors 0.04 -3.07 -0.199 -0.769 0.0764 0.467 -0.455 -0.195 -0.248 0.132 -0.329
mytilus_sp 0.06 -1.39 -0.0402 -0.0698 -0.0902 0.176 -0.269 -0.45 -0.0716 0.0639 -0.196
nematoda 0.33 -0.518 0.3 -0.576 0.0079 0.0241 -0.274 0.162 -0.248 -0.394 0.049
nemertea 0.11 -2.51 0.164 -0.608 0.223 0.281 -0.296 -0.148 -0.13 -0.043 0.278
neoleanira_tetragona 0.02 -2.88 -0.263 -0.537 -0.00709 0.173 -0.37 -0.193 -0.0584 0.00288 -0.243
nephtyidae_spp 0.03 -1.53 0.214 -0.195 -0.196 0.0406 -0.197 0.0525 0.000945 -0.154 0.0351
nephtys_caeca 0.1 -1.75 -0.467 -0.148 0.0994 0.0436 -0.348 -0.165 0.145 0.0271 -0.351
nephtys_incisa 0.28 -0.782 0.671 0.126 0.00538 0.0489 0.24 -0.104 -0.0849 -0.239 0.474
nephtys_sp 0.01 -2.31 0.214 -0.458 -0.029 0.318 -0.258 -0.0925 -0.11 0.0394 -0.0458
nuculana_minuta 0.07 -1.58 0.402 -0.376 0.0737 0.362 -0.219 -0.12 -0.156 -0.105 0.0925
nymphonidae_spp 0.07 -3.21 -0.192 -0.756 0.166 0.402 -0.445 -0.236 -0.3 0.078 -0.361
oenopota_sp 0.11 -1.95 -0.197 -0.429 0.0645 0.551 -0.396 -0.121 -0.345 -0.0331 -0.197
oligochaeta 0.09 -1.46 -0.216 0.0214 -0.123 0.274 -0.279 -0.23 0.0759 0.19 0.114
ophelia_limacina 0.02 -1.86 0.216 -0.366 0.1 0.204 -0.241 -0.108 -0.0678 0.0662 0.11
opheliidae_spp 0.08 -2.64 0.333 -0.562 0.222 0.391 -0.253 -0.0985 -0.199 -0.0772 -0.152
ophiopholis_aculeata 0.04 -2.71 0.0206 -0.648 0.0314 0.503 -0.363 -0.161 -0.301 -0.024 -0.273
ophiura_robusta 0.16 -2.65 -0.00219 -0.821 0.0748 0.35 -0.4 -0.166 -0.439 -0.085 -0.372
orchomenella_minuta 0.02 -2.14 -0.0218 -0.2 -0.0304 0.276 -0.263 -0.226 -0.238 0.014 -0.121
ostracoda 0.09 -0.684 0.24 -0.0661 -0.285 0.0606 0.166 -0.188 -0.145 -0.0262 0.0841
pagurus_sp 0.02 -2.84 -0.0676 -0.617 0.0584 0.447 -0.376 -0.227 -0.149 -0.027 -0.166
pagurus_pubescens 0.09 -2.77 0.112 -0.78 0.158 0.206 -0.333 -0.201 -0.276 -0.0504 -0.218
pandalus_montagui 0.07 -2.56 -0.0572 -0.372 0.0461 0.444 -0.31 -0.147 -0.153 0.139 0.164
parathyasira_equalis 0 -2.69 -0.0444 -0.489 0.00155 0.307 -0.313 -0.172 -0.14 -0.0143 -0.201
parvicardium_pinnulatum 0.03 -2.22 -0.000665 -0.62 0.0132 0.345 -0.308 -0.2 -0.138 0.0492 -0.194
periploma_leanum 0.03 -2.09 0.0717 -0.355 -0.0328 0.341 -0.277 -0.0149 -0.0175 0.0727 0.161
philine_lima 0.02 -2.82 -0.0575 -0.616 0.0503 0.441 -0.364 -0.226 -0.144 -0.0329 -0.168
philomedes_sp 0.15 -2.6 0.509 -0.748 0.145 0.194 -0.233 -0.064 -0.282 -0.099 -0.115
pholoe_longa 0.02 -2.84 -0.32 -0.532 0.0535 0.431 -0.423 -0.264 -0.115 0.247 -0.279
pholoe_sp 0.08 -0.0448 -0.0285 0.344 -0.0806 0.0277 0.0781 -0.179 -0.193 -0.0716 0.0199
phoxocephalus_holbolli 0.09 -0.958 -0.259 -0.00358 -0.0862 -0.0979 -0.0645 -0.206 0.155 -0.0627 -0.105
polynoidae_spp 0.07 -0.841 0.0755 0.152 -0.129 0.199 -0.213 -0.227 -0.0564 -0.107 0.128
pontogeneia_inermis 0.02 -2.79 -0.28 -0.492 0.0392 0.397 -0.404 -0.298 -0.112 0.302 -0.228
pontoporeia_femorata 0.32 -0.807 -0.173 0.23 -0.0165 0.248 -0.267 -0.123 0.0834 0.143 0.369
praxillella_praetermissa 0.07 -1.6 0.088 -0.118 -0.0662 0.394 -0.252 0.0547 0.0257 0.0946 0.0128
propebela_turricula 0.14 -2.06 0.643 -0.475 -0.101 0.254 -0.173 -0.039 -0.105 -0.0246 0.212
protomedeia_fasciata 0.07 -1.31 0.0288 0.0805 0.0373 0.119 -0.202 -0.271 0.158 -0.097 0.203
protomedeia_grandimana 0.17 -0.24 0.287 0.316 -0.0448 -0.0439 0.106 0.0414 -0.113 -0.071 0.33
puncturella_noachina 0.1 -2.9 -0.0512 -0.724 0.31 0.527 -0.408 -0.127 -0.361 0.00145 -0.336
quasimelita_formosa 0.1 -0.839 0.301 -0.0337 -0.0568 0.261 -0.2 -0.117 -0.0565 -0.0709 -0.0352
quasimelita_quadrispinosa 0 -2.61 -0.227 -0.335 0.0293 0.343 -0.355 -0.29 -0.129 0.233 -0.192
retusa_obtusa 0.08 -1.93 -0.502 -0.105 -0.0498 0.256 -0.318 -0.0382 0.132 0.246 -0.338
sabellidae_spp 0.1 -1.6 -0.332 -0.223 -0.0864 0.257 -0.0635 -0.166 0.258 0.037 -0.0475
scoletoma_fragilis 0.13 -1.6 -0.407 0.123 -0.0107 0.501 -0.0461 -0.191 -0.153 0.159 -0.179
scoletoma_sp 0.04 -2.78 -0.234 -0.704 0.0356 0.428 -0.441 -0.186 -0.252 0.1 -0.351
scoloplos_sp 0.01 -2.69 0.13 -0.534 0.0105 0.34 -0.302 -0.158 -0.143 -0.0126 -0.147
serripes_groenlandicus 0.11 -2.31 -0.0867 -0.135 0.298 0.318 -0.266 -0.0923 -0.0146 0.113 0.0973
sipuncula 0.16 -1.15 0.179 0.132 -0.154 0.248 0.202 0.16 -0.0345 -0.0445 -0.0333
solamen_glandula 0.08 -2.97 -0.0887 -0.634 0.0977 0.572 -0.404 -0.174 -0.316 0.0358 -0.35
solariella_sp 0.16 -2.35 0.308 -0.662 0.179 0.44 -0.282 -0.11 -0.387 -0.197 -0.225
strongylocentrotus_sp 0.14 -2.92 -0.109 -0.853 0.287 0.594 -0.456 -0.113 -0.433 -0.0194 -0.416
tachyrhynchus_erosus 0.01 -2.7 0.14 -0.525 0.00666 0.341 -0.295 -0.161 -0.139 -0.0229 -0.145
thracia_septentrionalis 0.14 -1.11 -0.482 -0.169 0.0908 0.255 -0.197 -0.171 -0.0538 -0.136 -0.181
thyasira_gouldi 0.17 -0.79 0.516 -0.161 -0.0686 0.217 -0.162 -0.0273 -0.195 -0.153 0.217
thyasira_sp 0.01 -2.7 0.136 -0.559 0.00787 0.346 -0.305 -0.156 -0.156 -0.00147 -0.141
trichotropis_bicarinata 0.01 -2.75 0.217 -0.582 -0.0103 0.312 -0.285 -0.157 -0.182 0.0491 -0.154
turritellopsis_stimpsoni 0.03 -2.32 0.288 -0.493 -0.00673 0.166 -0.225 -0.156 -0.227 -0.0712 0.0126
yoldia_myalis 0.01 -2.04 0.131 -0.275 0.0786 0.188 -0.246 -0.0794 -0.0791 -0.0197 0.0303
AVERAGED 0.09 -2.06 -0.0127 -0.371 0.0211 0.276 -0.256 -0.149 -0.133 0.009 -0.114
Ancillary information

Predictive power plot:

95 % confidence interval of the predictor coefficients estimated by the HMSC model:

Variance partitioning:

Trace plots:

Density plots:

Exposure indices
Model outcomes

Mean of the predictor coefficients estimated by the HMSC model:

Tjur’s pseudo-R2 and variables coefficients for each taxon
  Pseudo.R2 Intercept depth aquaculture dredging runoff sewers structures shipping fisheries
aceroides_aceroides_latipes 0.05 -0.967 0.00811 0.132 -0.322 0.0469 0.00115 0.0432 0.243 0.00976
akanthophoreus_gracilis 0.21 -0.866 0.353 0.287 -0.174 0.149 -0.129 -0.21 0.278 0.154
ameritella_agilis 0.04 -1.56 -0.0484 -0.0449 -0.143 -0.115 -0.127 0.0245 -0.238 0.0168
ameroculodes_edwardsi 0.13 -1.99 0.308 0.0534 -0.227 0.00925 -0.489 0.216 -0.0687 0.0172
ampelisca_vadorum 0.04 -2.89 -0.158 -0.332 -0.384 -0.528 -0.613 0.737 -0.0638 0.0857
amphipoda 0.05 -0.408 0.0847 0.0679 0.0991 0.188 0.13 -0.453 0.00346 0.0271
anonyx_lilljeborgi 0.06 -1.71 -0.298 -0.0409 -0.356 -0.341 -0.342 0.503 -0.0116 -0.167
anthozoa 0.07 -2.98 -0.305 -0.379 -0.418 -0.602 -0.693 0.854 -0.166 -0.0832
arcteobia_anticostiensis 0.04 -2.78 -0.198 0.0522 -0.387 -0.405 -0.511 0.719 -0.0977 -0.0156
arrhoges_occidentalis 0.03 -2.78 -0.148 -0.316 -0.397 -0.451 -0.647 0.696 -0.02 -0.0485
astarte_sp 0.09 -2.21 0.107 -0.296 -0.409 0.0248 -0.62 0.335 -0.0645 0.0514
axinopsida_orbiculata 0.11 -0.861 0.197 -0.0493 0.162 0.0871 -0.196 -0.0754 -0.0139 0.167
axiothella_catenata 0.02 -2.29 0.127 -0.124 -0.346 -0.405 -0.223 0.395 0.145 0.00511
bathymedon_longimanus 0.1 -2.2 -0.345 0.173 -0.127 -0.214 -0.499 0.716 -0.124 -0.026
bathymedon_obtusifrons 0.02 -2.4 -0.25 -0.193 -0.447 -0.348 -0.158 0.694 -0.124 -0.117
bipalponephtys_neotena 0.13 1.08 0.388 0.274 0.127 0.394 0.504 -0.584 0.439 0.0192
boreochiton_ruber 0.12 -2.42 -0.395 -0.218 -0.33 -0.463 -0.732 0.819 -0.205 0.159
brachydiastylis_sp 0.1 -2.7 -0.0254 -0.307 -0.399 -0.409 -0.72 0.623 0.0527 0.118
byblis_gaimardii 0.19 -2.47 0.0254 -0.211 -0.359 -0.167 -0.774 0.416 -0.0253 0.238
cancer_irroratus 0.01 -2.67 -0.352 -0.272 -0.361 -0.286 -0.302 0.535 -0.258 -0.0757
caprella_septentrionalis 0.17 -2.26 -0.639 -0.319 -0.391 -0.037 -0.236 0.433 -0.465 -0.155
chaetodermatida 0.14 -1.12 0.451 0.0965 -0.178 0.4 -0.0375 -0.176 0.069 0.135
chionoecetes_opilio 0.04 -2.43 0.176 -0.0373 -0.388 -0.292 -0.471 0.513 0.0249 -0.0757
chlamys_islandica 0.27 -3.08 -0.332 -0.322 -0.373 -0.502 -0.608 0.807 -0.177 0.259
chone_sp 0.02 -2.8 -0.383 -0.255 -0.443 -0.298 -0.441 0.767 -0.222 0.0163
ciliatocardium_ciliatum 0.03 -2.78 9.11e-05 -0.296 -0.395 -0.487 -0.508 0.66 -0.0778 0.0545
cirripedia 0.03 -2.89 -0.457 -0.321 -0.415 -0.571 -0.455 0.915 -0.196 -0.0511
cistenides_granulata 0.14 -1.08 -0.022 0.0502 -0.194 -0.108 -0.419 0.332 -0.00262 0.176
cossura_longocirrata 0.01 -2.52 -0.142 -0.213 -0.356 -0.465 -0.309 0.527 0.179 -0.0718
crassicorophium_bonellii 0.02 -2.4 -0.212 -0.294 -0.316 -0.446 -0.506 0.691 0.00679 -0.00587
crenella_decussata 0.13 -1.87 -0.148 -0.0598 -0.369 -0.416 -0.714 0.72 -0.0588 0.0273
cumacea 0.03 -2.24 0.219 -0.148 -0.242 -0.456 0.0248 0.491 0.0106 -0.0989
cyclocardia_borealis 0.07 -2.96 -0.294 -0.377 -0.412 -0.577 -0.686 0.827 -0.179 -0.0802
cyrtodaria_siliqua 0.02 -2.58 -0.0771 -0.177 -0.152 -0.263 -0.391 0.731 -0.0814 -0.0387
diastylis_rathkei 0.09 -1.76 -0.392 -0.342 0.0858 -0.203 -0.602 0.532 -0.0973 -0.0589
diastylis_sculpta 0.14 -2.12 -0.503 -0.257 -0.438 -0.253 -0.546 0.929 -0.0415 -0.0899
diastylis_sp 0.03 -2.71 0.139 -0.159 -0.387 -0.402 -0.415 0.611 -0.0848 0.00252
echinarachnius_parma 0.13 -1.36 -0.215 0.0662 -0.18 -0.00403 -0.56 0.627 -0.0499 0.091
edotia_montosa 0.07 -2.53 -0.621 -0.332 -0.283 -0.452 -0.412 0.87 -0.302 -0.158
ennucula_tenuis 0.33 -0.221 0.713 0.38 -0.0152 0.376 -0.0132 -0.262 0.426 0.139
eteone_sp 0.1 -1.96 0.115 -0.253 -0.318 -0.455 -0.101 0.295 0.137 0.18
euchone_sp 0.02 -2.74 -0.333 -0.25 -0.449 -0.32 -0.401 0.762 -0.15 -0.0661
eudorella_emarginata 0.12 -1.86 -0.0592 -0.169 -0.194 -0.575 0.0792 0.356 0.308 -0.159
eudorellopsis_integra 0.3 0.235 0.4 0.107 0.12 0.0614 0.139 -0.401 0.659 -0.0982
euspira_pallida 0.02 -2.57 -0.00542 -0.224 -0.388 -0.1 -0.523 0.432 -0.0298 -0.0193
glycera_capitata 0.02 -2.44 -0.282 -0.324 -0.361 -0.591 -0.359 0.692 -0.0897 0.0126
glycera_sp 0.1 -2.21 -0.545 -0.404 -0.339 -0.43 -0.361 0.49 -0.332 -0.221
goniada_maculata 0.3 -0.362 0.643 0.272 -0.192 0.182 0.142 -0.276 0.308 0.122
guernea_prinassus_nordenskioldi 0.01 -2.36 -0.178 -0.251 -0.399 -0.434 -0.226 0.697 -0.135 -0.0633
halacaridae_spp 0.26 -3.08 -0.336 -0.313 -0.388 -0.504 -0.588 0.786 -0.173 0.252
haminoea_solitaria 0.04 -2.76 -0.175 0.0303 -0.373 -0.4 -0.491 0.698 -0.12 -0.00272
hardametopa_carinata 0.02 -2.74 -0.34 -0.251 -0.437 -0.318 -0.392 0.744 -0.171 -0.07
harmothoe_sp 0.06 -2.29 0.113 -0.299 -0.351 -0.49 -0.372 0.492 0.029 0.0832
harpacticoida 0.2 -0.194 -0.236 -0.146 -0.319 0.504 -0.0611 -0.267 0.113 0.0344
hediste_diversicolor 0.2 -1.57 -0.682 -0.311 -0.0219 -0.382 0.0346 0.559 -0.323 -0.267
heteranomia_squamula 0.02 -2.82 -0.325 -0.263 -0.389 -0.366 -0.529 0.897 -0.141 -0.0394
hiatella_arctica 0.05 -2.3 -0.125 -0.227 -0.456 -0.291 -0.36 0.844 -0.103 -0.0816
holothuroidea 0.1 -2.47 -0.239 -0.263 -0.355 0.143 -0.748 0.493 -0.141 0.111
idotea_phosphorea 0.09 -2.45 -0.5 -0.303 -0.355 -0.0722 -0.217 0.334 -0.431 -0.122
ischyroceridae_spp 0.02 -2.7 -0.265 -0.203 -0.447 -0.305 -0.409 0.811 -0.103 -0.0679
ischyrocerus_anguipes 0.14 -2.41 -0.703 -0.4 -0.379 -0.3 -0.383 0.621 -0.459 -0.182
isopoda 0.09 -2.89 -0.168 -0.279 -0.375 -0.157 -0.655 0.53 -0.143 0.193
lacuna_vincta 0.05 -2.58 -0.506 -0.304 -0.395 -0.299 -0.375 0.68 -0.289 -0.0794
lamprops_fuscatus 0.06 -1.33 -0.124 0.167 -0.117 0.0513 -0.206 0.372 0.0767 0.0738
lamprops_quadriplicata 0.04 -2.53 -0.494 -0.297 -0.324 -0.451 -0.22 0.686 -0.263 -0.147
lepeta_caeca 0.11 -2 -0.189 -0.254 -0.362 -0.273 -0.505 0.461 -0.256 0.128
leucon_leucon_nasicoides 0.29 -0.341 0.577 0.237 -0.0842 -0.192 0.335 -0.0156 0.353 -0.0644
littorina_littorea 0.08 -2.25 -0.384 -0.272 -0.38 -0.0565 -0.204 0.367 -0.434 -0.154
lumbrineridae_spp 0.01 -2.35 -0.0364 -0.184 -0.38 -0.42 -0.346 0.645 -0.125 -0.0283
lysianassidae_spp 0.05 -2.61 -0.32 0.0324 -0.342 -0.295 -0.584 0.851 -0.141 0.0231
macoma_calcarea 0.06 0.791 0.152 0.117 0.131 0.682 0.00347 -0.597 0.0765 0.0991
maera_danae 0.02 -2.81 -0.377 -0.276 -0.417 -0.287 -0.427 0.731 -0.234 0.0141
maldane_sarsi 0.03 -2.26 0.183 -0.135 -0.324 -0.404 0.0223 0.528 0.0248 0.0441
maldanidae_spp 0.18 -0.882 0.417 0.216 -0.269 -0.0672 0.288 0.0741 0.303 -0.09
monoculopsis_longicornis 0.16 -2.37 -0.0163 -0.241 -0.377 0.276 -0.664 0.179 -0.0738 0.11
muculus_musculus_discors 0.25 -3.07 -0.337 -0.331 -0.378 -0.503 -0.594 0.789 -0.177 0.246
mytilus_sp 0.09 -1.4 0.0679 0.0446 -0.282 0.212 -0.161 0.39 -0.292 -0.0627
nematoda 0.22 -0.353 0.205 0.126 -0.361 0.304 -0.18 -0.326 -0.0119 0.0726
nemertea 0.03 -2.3 -0.0688 -0.234 -0.319 -0.498 -0.288 0.46 0.194 0.0295
neoleanira_tetragona 0.01 -2.68 -0.387 -0.281 -0.371 -0.289 -0.311 0.54 -0.279 -0.0968
nephtyidae_spp 0.15 -1.62 0.0454 -0.0137 0.159 0.0928 -0.248 -0.127 -0.153 0.251
nephtys_caeca 0.07 -1.78 -0.528 -0.221 -0.176 -0.254 -0.496 0.582 -0.191 -0.0396
nephtys_incisa 0.23 -0.79 0.654 0.19 -0.172 0.000547 0.107 -0.0663 0.309 -0.0276
nephtys_sp 0.02 -2.24 0.136 -0.177 -0.372 -0.319 -0.215 0.376 0.17 -0.0974
nuculana_minuta 0.11 -1.52 0.347 0.0515 -0.402 -0.239 0.00338 0.272 0.142 0.166
nymphonidae_spp 0.02 -2.81 -0.294 -0.243 -0.39 -0.386 -0.497 0.895 -0.123 -0.0496
oenopota_sp 0.05 -1.77 -0.171 -0.289 -0.312 -0.0618 -0.411 0.291 -0.0163 0.0413
oligochaeta 0.09 -1.41 -0.176 -0.126 -0.196 -0.139 0.093 0.188 0.271 -0.11
ophelia_limacina 0.05 -1.87 0.174 -0.19 -0.399 -0.253 -0.183 0.306 0.32 -0.0256
opheliidae_spp 0.06 -2.57 0.127 -0.234 -0.38 -0.432 -0.531 0.581 -0.031 0.0757
ophiopholis_aculeata 0.11 -2.59 -0.0763 -0.0303 -0.347 -0.363 -0.486 0.567 -0.065 0.221
ophiura_robusta 0.07 -2.06 -0.176 -0.203 -0.371 -0.362 -0.593 0.735 -0.107 0.0912
orchomenella_minuta 0.02 -2.09 0.0263 -0.179 -0.438 -0.0605 -0.238 0.438 -0.0729 -0.0956
ostracoda 0.1 -0.642 0.246 0.0987 -0.232 0.377 -0.198 -0.164 0.252 0.0416
pagurus_sp 0.04 -2.79 -0.196 0.046 -0.387 -0.424 -0.513 0.732 -0.109 -0.0116
pagurus_pubescens 0.03 -2.22 -0.0514 -0.0338 -0.418 -0.258 -0.411 0.742 -0.121 -0.0644
pandalus_montagui 0.01 -2.55 -0.25 -0.234 -0.0361 -0.555 -0.29 0.695 -0.00448 -0.0788
parathyasira_equalis 0.04 -2.77 -0.186 0.0485 -0.392 -0.38 -0.5 0.703 -0.126 -0.0022
parvicardium_pinnulatum 0.03 -2.09 -0.122 -0.029 -0.45 -0.245 -0.403 0.636 -0.154 -0.052
periploma_leanum 0.09 -2.15 0.049 -0.0955 -0.336 -0.403 0.0355 0.32 0.335 -0.0887
philine_lima 0.04 -2.75 -0.188 0.0475 -0.382 -0.392 -0.497 0.696 -0.107 -0.0204
philomedes_sp 0.14 -2.23 0.205 -0.301 -0.231 -0.256 -0.635 0.431 0.111 0.107
pholoe_longa 0.02 -2.73 -0.335 -0.249 -0.456 -0.315 -0.402 0.76 -0.161 -0.0716
pholoe_sp 0.06 -0.069 0.0779 0.107 -0.0857 0.163 -0.232 0.0788 0.161 -0.129
phoxocephalus_holbolli 0.11 -1.02 -0.255 -0.0673 -0.17 0.119 -0.13 0.0714 -0.336 0.0189
polynoidae_spp 0.08 -0.843 0.227 -0.102 -0.357 0.0184 0.0913 -0.0661 0.00283 -0.199
pontogeneia_inermis 0.02 -2.79 -0.339 -0.245 -0.468 -0.31 -0.458 0.817 -0.147 -0.0699
pontoporeia_femorata 0.17 -0.813 -0.138 -0.172 -0.00876 -0.275 0.413 0.166 0.0456 -0.255
praxillella_praetermissa 0.08 -1.58 0.117 0.175 -0.0334 -0.393 0.178 0.416 0.256 -0.127
propebela_turricula 0.14 -2.03 0.473 -0.126 -0.337 -0.292 -0.0854 0.199 0.227 0.0971
protomedeia_fasciata 0.03 -1.29 -0.0715 -0.123 0.107 -0.192 -0.219 0.215 0.00771 0.0707
protomedeia_grandimana 0.13 -0.247 0.42 0.115 0.0515 0.185 0.378 -0.215 0.0697 -0.119
puncturella_noachina 0.09 -2.69 -0.206 -0.398 -0.396 -0.607 -0.703 0.758 -0.0666 0.0127
quasimelita_formosa 0.14 -0.781 0.402 0.357 -0.149 0.295 0.2 -0.138 0.072 0.057
quasimelita_quadrispinosa 0.02 -2.72 -0.259 -0.221 -0.437 -0.301 -0.415 0.797 -0.103 -0.0625
retusa_obtusa 0.11 -2.02 -0.501 -0.203 -0.0922 -0.341 -0.249 0.833 -0.0469 -0.0597
sabellidae_spp 0.1 -1.65 -0.448 -0.3 -0.354 -0.528 -0.0545 0.529 -0.0252 -0.134
scoletoma_fragilis 0.11 -1.72 -0.389 -0.343 -0.236 -0.693 -0.12 0.859 -0.143 -0.268
scoletoma_sp 0.16 -2.58 -0.363 -0.305 -0.000977 -0.345 -0.552 0.617 -0.156 0.229
scoloplos_sp 0.03 -2.73 -0.0405 0.00716 -0.38 -0.385 -0.45 0.657 -0.108 0.058
serripes_groenlandicus 0.03 -2.24 -0.0793 -0.153 -0.138 -0.506 -0.157 0.507 0.27 -0.0868
sipuncula 0.06 -1.11 0.212 0.0428 -0.0931 -0.266 0.134 0.239 0.0989 -0.168
solamen_glandula 0.07 -3 -0.298 -0.38 -0.414 -0.591 -0.689 0.83 -0.167 -0.0825
solariella_sp 0.15 -2.07 0.107 -0.11 -0.34 -0.241 -0.614 0.537 0.0652 0.242
strongylocentrotus_sp 0.23 -2.75 -0.303 -0.44 -0.365 -0.598 -0.768 0.733 -0.142 0.216
tachyrhynchus_erosus 0.03 -2.73 -0.0427 0.0197 -0.377 -0.365 -0.436 0.643 -0.12 0.038
thracia_septentrionalis 0.08 -1.03 -0.438 -0.0716 -0.151 -0.135 -0.234 0.21 0.0265 0.013
thyasira_gouldi 0.22 -0.8 0.554 0.195 -0.384 0.198 0.0139 -0.145 0.368 0.0831
thyasira_sp 0.03 -2.72 -0.0476 0.0193 -0.385 -0.368 -0.442 0.666 -0.117 0.0463
trichotropis_bicarinata 0 -2.5 0.0519 -0.193 -0.229 -0.197 -0.448 0.534 -0.0038 -0.0424
turritellopsis_stimpsoni 0.04 -2.31 0.0573 -0.159 -0.375 -0.108 -0.549 0.358 -0.118 0.0385
yoldia_myalis 0.02 -2.09 0.0776 0.0101 -0.405 -0.363 -0.205 0.532 0.0911 -0.0852
AVERAGED 0.09 -1.98 -0.0874 -0.128 -0.284 -0.229 -0.308 0.441 -0.0324 -0.00197
Ancillary information

Predictive power plot:

95 % confidence interval of the predictor coefficients estimated by the HMSC model:

Variance partitioning:

Trace plots:

Density plots:

2.2.2. Density data

We considered density data with an overPoisson distribution. Diagnostics are here.

All variables
Model outcomes

Mean of the predictor coefficients estimated by the HMSC model:

Nakagawa’s pseudo-R2 and variables coefficients for each taxon
  Pseudo.R2 Intercept depth om gravel silt clay arsenic cadmium copper mercury aquaculture dredging runoff sewers structures shipping fisheries
aceroides_aceroides_latipes 0.22 -1.52 0.0109 -0.148 -0.399 0.198 -0.38 -0.443 -0.156 0.294 -0.0157 0.316 -0.482 -0.061 0.226 -0.103 0.39 0.0099
akanthophoreus_gracilis 0.12 -1.44 0.573 -0.188 -0.315 -0.0345 0.314 0.17 -0.64 0.101 0.0577 0.558 -0.378 0.665 -0.0347 -0.947 0.55 0.374
ameritella_agilis 0.24 -2.07 0.0183 0.192 -0.0818 -0.463 0.221 -0.271 0.175 0.141 -0.0972 0.0412 -0.101 0.22 -0.176 -0.556 -0.213 0.0142
ameroculodes_edwardsi 0.37 -2.75 0.65 -0.247 0.0711 0.0306 -0.214 -0.358 -0.233 0.0675 0.106 0.14 0.00781 0.324 -0.338 -0.261 -0.145 -0.0274
ampelisca_vadorum 0.41 -4.44 -0.0442 -0.835 0.303 0.373 -0.515 -0.585 -0.704 0.411 -0.196 -0.389 -0.585 -0.746 -0.137 0.559 -0.28 -0.0571
amphipoda 0.04 -0.611 0.135 0.136 0.0742 0.0336 -0.207 -0.0372 0.365 -0.348 0.231 0.105 0.129 0.56 0.0198 -0.657 -0.0283 0.0936
anonyx_lilljeborgi 0.37 -2.54 -0.604 -0.208 0.0274 0.189 -0.485 -0.721 -0.482 0.47 -0.256 0.00403 -0.707 -0.754 0.0371 0.394 -0.0215 -0.39
anthozoa 0.38 -4.3 -0.31 -0.709 0.0481 0.645 -0.586 -0.619 -0.736 0.416 -0.26 -0.45 -0.656 -0.938 -0.196 0.899 -0.356 -0.198
arcteobia_anticostiensis 0.37 -3.91 -0.014 -0.751 -0.137 0.354 -0.481 -0.854 -0.283 0.501 -0.244 0.241 -0.338 -0.601 0.0121 0.486 -0.212 -0.092
arrhoges_occidentalis 0.37 -4.09 0.031 -0.72 -0.0951 0.181 -0.421 -0.644 -0.527 0.477 -0.239 -0.302 -0.484 -0.576 -0.308 0.373 -0.0475 -0.153
astarte_sp 0.48 -3.39 0.416 -0.571 -0.0628 0.167 -0.372 -0.249 -0.88 0.0903 -0.0703 -0.313 -0.629 0.126 -0.109 -0.0743 -0.249 -0.0283
axinopsida_orbiculata 0.1 -1.41 0.462 0.113 -0.346 -0.245 0.176 -0.166 0.695 -0.156 -0.389 0.0489 0.518 0.332 -0.515 -0.148 0.207 0.379
axiothella_catenata 0.26 -3.11 0.417 -0.228 -0.0933 0.165 -0.303 -0.472 -0.219 -0.0085 0.694 -0.19 -0.454 -0.301 -0.143 0.251 0.0305 -0.00102
bathymedon_longimanus 0.42 -3.21 -0.352 -0.548 -0.253 0.238 -0.445 -0.995 0.0423 0.442 -0.51 0.476 0.067 -0.51 -0.143 0.688 -0.158 -0.0871
bathymedon_obtusifrons 0.42 -4.18 -0.47 -1.19 -0.0828 0.878 -0.958 -0.87 -0.757 0.947 -0.576 -0.141 -1.01 -1.18 0.989 0.968 -0.493 -0.36
bipalponephtys_neotena 0 1.85 -0.157 0.662 -0.119 0.305 0.0397 -0.0233 0.345 -0.706 0.268 0.38 -0.431 0.556 -0.138 0.118 0.55 0.203
boreochiton_ruber 0.55 -4.63 -0.523 -1.43 0.239 0.865 -0.819 -0.758 -1.19 0.25 -0.507 -0.301 -0.65 -1.18 0.00682 1.32 -0.74 0.084
brachydiastylis_sp 0.39 -3.85 0.243 -0.66 0.149 0.244 -0.335 -0.398 -0.541 0.258 -0.136 -0.293 -0.434 -0.416 -0.469 0.337 0.108 0.079
byblis_gaimardii 0.44 -3.51 0.412 -0.43 -0.033 -0.0526 -0.222 -0.434 -0.36 0.135 0.0662 -0.121 -0.23 -0.103 -0.676 0.0442 -0.0126 0.211
cancer_irroratus 0.43 -4.22 -0.445 -0.822 0.00838 0.274 -0.702 -0.729 -0.488 0.48 -0.247 -0.328 -0.561 -0.505 0.406 0.282 -0.713 -0.252
caprella_septentrionalis 0.05 -5.65 -1.82 -1.08 -0.0155 0.012 -0.795 -1.54 -0.765 0.8 -0.501 -0.908 -1.06 -1.06 0.705 0.699 -1.64 -0.715
chaetodermatida 0.35 -1.57 0.745 -0.155 -0.358 -0.298 0.175 -0.274 -0.186 0.0278 0.0471 0.267 -0.137 0.579 0.355 -0.618 0.105 0.155
chionoecetes_opilio 0.36 -3.42 0.483 -0.444 -0.0904 0.403 -0.326 -0.464 -0.669 0.315 -0.182 0.0961 -0.519 -0.301 -0.0484 0.232 0.0509 -0.103
chlamys_islandica 0.39 -4.34 -0.183 -0.874 -0.0898 0.46 -0.572 -0.691 -0.526 0.511 -0.308 -0.27 -0.435 -0.77 -0.0143 0.632 -0.324 0.164
chone_sp 0.38 -4.56 -0.704 -0.669 -0.122 0.409 -0.749 -0.977 -0.354 0.755 -0.261 -0.373 -0.687 -0.595 0.133 0.782 -0.681 -0.0924
ciliatocardium_ciliatum 0.33 -3.98 0.363 -0.561 -0.0157 -0.0134 -0.381 -0.573 -0.443 0.414 -0.0337 -0.261 -0.393 -0.485 -0.102 0.222 -0.234 -0.027
cirripedia 0.31 -3.91 -0.521 -0.5 -0.0793 0.331 -0.51 -0.79 0.456 0.112 -0.223 -0.412 -0.297 -0.883 -0.288 1.15 -0.249 -0.106
cistenides_granulata 0.33 -1.63 -0.0841 -0.307 0.0611 0.334 -0.381 -0.161 -0.453 -0.431 -0.00407 -0.0153 -0.368 -0.404 -0.123 0.661 -0.0736 0.109
cossura_longocirrata 0.19 -3.44 -0.211 -0.0889 -0.0778 0.383 -0.283 -0.341 0.105 0.416 -0.105 -0.231 -0.443 -0.45 -0.507 0.393 0.637 -0.0753
crassicorophium_bonellii 0.28 -3.53 -0.00217 -0.809 -0.13 0.00108 -0.387 -0.046 -0.409 0.488 -0.247 -0.253 -0.399 -0.445 -0.352 0.408 0.0965 -0.0249
crenella_decussata 0.63 -3.68 0.0721 -1.27 0.298 0.458 -0.551 -0.63 -1.11 0.362 -0.471 0.135 -0.636 -0.745 0.242 0.548 -0.422 -0.129
cumacea 0.21 -3.13 0.702 -0.239 -0.257 -0.199 0.307 -0.302 -0.248 0.516 -0.0464 0.0822 -0.339 -0.135 0.189 -0.173 0.15 -0.0824
cyclocardia_borealis 0.44 -4.47 -0.387 -0.794 0.119 0.82 -0.64 -0.653 -0.939 0.421 -0.326 -0.563 -0.784 -1.14 -0.273 1.08 -0.434 -0.252
cyrtodaria_siliqua 0.32 -3.78 0.216 -0.629 -0.12 0.274 -0.428 -0.59 -0.437 0.366 -0.26 -0.0707 -0.0315 -0.295 0.0145 0.544 -0.256 -0.0509
diastylis_rathkei 0.15 -2.97 -0.906 0.0111 0.26 0.326 -0.381 -0.35 -0.0965 0.066 0.206 -0.868 0.0994 -0.456 -1.34 1.15 -0.199 -0.154
diastylis_sculpta 0.37 -3.41 -1.09 -0.277 -0.139 0.636 -0.539 -0.822 -0.11 0.594 -0.259 -0.616 -0.748 -1.01 -0.965 1.8 0.0278 -0.26
diastylis_sp 0.25 -3.52 0.487 -0.309 -0.0784 0.365 -0.321 -0.454 -0.267 0.211 -0.0256 -0.0843 -0.383 -0.353 -0.165 0.445 -0.112 0.00918
echinarachnius_parma 0.56 -2.82 -0.193 -1.5 -0.249 0.465 -0.676 -0.9 -0.859 0.663 -1.04 0.401 -0.188 -0.53 0.45 0.543 -0.161 -0.0901
edotia_montosa 0.49 -4.12 -0.907 -0.687 -0.18 -0.166 -0.669 -0.765 -0.351 0.9 -0.576 -0.409 -0.377 -0.767 0.115 0.605 -0.74 -0.414
ennucula_tenuis 0.16 -0.303 0.871 0.123 0.00743 0.153 0.252 0.221 -0.342 -0.495 0.255 0.482 -0.143 0.636 -0.111 -0.269 0.558 0.306
eteone_sp 0.47 -3.09 0.303 -0.949 0.0729 0.262 -0.521 -0.406 -0.796 0.528 -0.354 -0.0323 -0.624 -0.566 0.688 -0.258 0.179 0.082
euchone_sp 0.22 -4.98 -0.838 -0.772 -0.132 0.605 -0.827 -1.25 -0.453 1.1 -0.409 -0.378 -0.901 -0.887 0.176 0.994 -0.471 -0.4
eudorella_emarginata 0.32 -2.53 -0.154 0.332 0.339 0.388 -0.269 -0.137 -0.151 0.00545 0.308 -0.329 -0.594 -0.376 -0.17 0.393 0.409 -0.147
eudorellopsis_integra 0.01 0.715 0.201 0.846 -0.0918 0.0459 0.15 0.941 0.332 -1.42 0.767 -0.0232 -0.41 0.911 -0.715 -0.0332 1.24 0.218
euspira_pallida 0.29 -3.61 0.278 -0.472 -0.0839 0.194 -0.345 -0.394 -0.545 0.366 -0.0503 -0.14 -0.45 0.0689 -0.199 -0.0508 0.00213 -0.0514
glycera_capitata 0.28 -3.64 -0.271 -0.763 0.32 0.466 0.0562 -0.522 -0.53 0.46 -0.251 -0.334 -0.604 -0.867 -0.0325 0.629 -0.112 -0.123
glycera_sp 0.4 -3.19 -0.95 0.0612 0.0649 0.36 -0.557 -0.614 0.0182 0.41 -0.0592 -0.696 -0.586 -0.668 -0.365 0.598 -0.494 -0.399
goniada_maculata 0.3 -0.54 0.741 0.116 -0.402 0.0501 -0.246 -0.354 0.167 -0.336 0.172 0.205 -0.136 0.186 0.226 -0.115 0.307 0.184
guernea_prinassus_nordenskioldi 0.41 -4.43 -0.291 -1.57 -0.0734 1.03 -1.01 -0.845 -0.975 0.909 -0.702 -0.161 -0.928 -1.41 1.01 1.08 -0.563 -0.277
halacaridae_spp 0.42 -4.49 -0.213 -0.958 -0.0621 0.499 -0.623 -0.705 -0.573 0.533 -0.33 -0.279 -0.491 -0.8 0.0173 0.662 -0.368 0.163
haminoea_solitaria 0.33 -3.69 0.0408 -0.562 -0.205 0.137 -0.408 -0.739 -0.295 0.552 -0.277 0.284 -0.334 -0.467 0.0232 0.302 -0.167 -0.0706
hardametopa_carinata 0.38 -4.13 -0.553 -0.608 -0.0725 0.453 -0.647 -0.887 -0.36 0.696 -0.283 -0.306 -0.677 -0.716 0.09 0.804 -0.341 -0.259
harmothoe_sp 0.37 -3.32 0.315 -0.53 0.257 0.597 -0.443 -0.312 -0.762 0.21 -0.156 -0.268 -0.656 -0.66 0.109 0.429 0.0167 0.043
harpacticoida 0.05 -0.253 -0.503 -0.0081 0.00158 -0.179 -0.338 0.611 0.0223 -0.631 -0.137 -0.201 -0.551 0.642 0.175 -0.235 0.125 -0.0299
hediste_diversicolor 0.48 -2.02 -1.01 0.264 0.161 0.652 -0.295 0.261 -0.0143 -0.644 0.48 -0.661 -0.336 -0.22 -0.207 1.12 -0.675 -0.219
heteranomia_squamula 0.49 -4.57 -0.396 -1 0.196 0.557 -0.72 -0.73 -0.811 0.328 -0.306 -0.384 -0.668 -0.84 0.0782 1.14 -0.51 -0.203
hiatella_arctica 0.54 -3.95 0.0677 -1 0.0584 0.251 -0.656 -0.68 -0.851 0.242 -0.261 -0.213 -0.779 -0.639 0.449 0.787 -0.554 -0.221
holothuroidea 0.4 -3.34 -0.0996 -0.244 -0.209 -0.101 -0.327 -0.485 -0.681 0.453 -0.123 -0.206 -0.402 0.427 -0.356 -0.202 -0.198 0.0201
idotea_phosphorea 0.49 -3.99 -0.815 -0.722 0.139 0.577 -0.898 -0.617 -0.67 0.266 -0.177 -0.484 -0.666 -0.285 0.589 0.162 -1.12 -0.342
ischyroceridae_spp 0.26 -3.65 -0.329 -0.236 -0.1 0.366 -0.464 -0.696 -0.306 0.495 -0.0874 -0.253 -0.622 -0.468 -0.117 0.812 -0.151 -0.181
ischyrocerus_anguipes 0.09 -5.54 -2.01 -0.283 -0.253 -0.233 -0.956 -1.82 0.0759 1.12 -0.447 -1.08 -0.839 -1.43 -0.204 1.18 -1.36 -0.77
isopoda 0.3 -3.65 0.147 -0.349 -0.154 -0.025 -0.3 -0.461 -0.285 0.271 -0.0177 -0.186 -0.263 0.104 -0.336 -0.0642 -0.137 0.168
lacuna_vincta 0.57 -4.77 -0.947 -1.07 0.0487 0.383 -0.883 -0.933 -0.794 0.439 -0.4 -0.521 -0.832 -0.968 0.41 0.898 -0.891 -0.331
lamprops_fuscatus 0.31 -2.13 -0.035 -0.576 -0.331 0.126 -0.39 -0.564 -0.197 0.444 -0.352 0.375 -0.0528 0.0546 0.229 0.0814 0.153 0.134
lamprops_quadriplicata 0.4 -3.95 -0.584 -0.781 -0.0892 -0.113 -0.655 -0.44 -0.369 0.537 -0.267 -0.318 -0.529 -0.623 0.413 0.332 -0.613 -0.306
lepeta_caeca 0.59 -3.63 -0.113 -1.09 0.365 0.58 -0.726 -0.59 -0.993 0.27 -0.285 -0.277 -0.695 -0.734 0.319 0.426 -0.823 -0.0768
leucon_leucon_nasicoides 0.01 -0.516 0.968 0.784 -0.152 -0.179 0.115 0.172 -0.0726 -0.433 0.487 0.24 -0.334 0.0598 0.243 0.078 0.548 0.231
littorina_littorea 0.39 -3.22 -0.601 -0.0807 -0.0341 0.308 -0.665 -0.554 -0.408 0.373 -0.166 -0.39 -0.621 -0.0878 0.381 0.103 -0.884 -0.314
lumbrineridae_spp 0.21 -3.19 0.268 -0.155 -0.132 0.351 -0.333 -0.546 0.734 -0.374 -0.0529 -0.307 -0.189 -0.614 -0.438 1.02 -0.142 0.0739
lysianassidae_spp 0.48 -4.09 -0.314 -0.826 -0.189 0.467 -0.503 -1.03 -0.694 0.911 -0.706 0.256 -0.382 -0.682 0.0256 0.796 -0.27 -0.0966
macoma_calcarea 0.01 0.999 -0.095 0.155 -0.063 0.0665 -0.224 0.0973 -0.0924 -0.482 -0.151 0.235 0.0568 0.96 0.28 -0.709 -0.000779 0.125
maera_danae 0.38 -4.09 -0.504 -0.594 -0.102 0.362 -0.626 -0.799 -0.357 0.603 -0.209 -0.307 -0.609 -0.535 0.109 0.64 -0.496 -0.12
maldane_sarsi 0.21 -2.99 0.39 -0.141 -0.0833 0.546 -0.389 0.134 -0.4 0.129 -0.0538 -0.0847 -0.586 -0.319 0.29 0.549 0.0116 0.132
maldanidae_spp 0.14 -1.56 0.706 0.332 -0.46 0.41 0.155 -0.132 -0.393 0.456 -0.132 0.536 -0.676 0.0728 0.241 0.0842 0.989 0.00502
monoculopsis_longicornis 0.36 -3.09 0.255 -0.193 -0.178 -0.134 -0.272 -0.172 -0.575 0.142 0.12 -0.181 -0.358 0.837 -0.374 -0.679 -0.0818 0.13
muculus_musculus_discors 0.41 -4.43 -0.196 -0.913 -0.0739 0.486 -0.612 -0.701 -0.552 0.531 -0.322 -0.289 -0.466 -0.79 -0.00295 0.642 -0.367 0.154
mytilus_sp 0.02 -2.54 -0.0742 -0.193 -0.141 0.302 -0.578 -0.856 -0.24 -0.0511 -0.146 -0.0345 -0.388 -0.273 0.202 0.924 -0.763 -0.15
nematoda 0.01 -1.19 0.531 -1.39 -0.0744 -0.158 -0.692 0.137 -1.32 0.124 -0.445 0.51 -0.821 0.386 1.21 -1.61 -0.141 0.0443
nemertea 0.29 -3.62 0.0631 -0.587 0.432 0.172 -0.391 -0.469 -0.37 0.0933 0.596 -0.482 -0.593 -0.593 -0.253 0.268 0.166 -0.0657
neoleanira_tetragona 0.42 -4.11 -0.366 -0.648 -0.137 -0.0584 -0.623 -0.772 -0.306 0.385 -0.187 -0.323 -0.459 -0.406 0.334 0.128 -0.658 -0.235
nephtyidae_spp 0.23 -2.09 0.31 -0.0667 -0.325 -0.218 -0.291 0.0878 0.238 -0.151 0.228 0.0273 0.422 0.507 -0.19 -0.596 -0.357 0.336
nephtys_caeca 0.28 -2.32 -0.628 -0.0171 0.151 -0.124 -0.311 -0.176 0.305 -0.0963 -0.15 -0.303 -0.0494 -0.156 -0.671 0.548 -0.11 -0.0619
nephtys_incisa 0.26 -1.1 0.748 0.309 -0.0164 0.00264 0.246 -0.157 -0.306 -0.109 0.304 0.249 -0.308 0.411 -0.155 -0.207 0.507 -0.0467
nephtys_sp 0.28 -3.09 0.379 -0.419 -0.159 0.231 -0.329 -0.469 -0.44 0.441 -0.185 -0.00873 -0.531 -0.268 0.207 -0.0657 0.348 -0.14
nuculana_minuta 0.44 -2.19 0.446 -0.504 0.0606 0.482 -0.426 -0.351 -0.551 0.033 -0.0625 0.161 -0.702 -0.463 0.465 0.322 0.209 0.112
nymphonidae_spp 0.47 -4.57 -0.323 -0.976 0.136 0.504 -0.684 -0.762 -0.781 0.405 -0.302 -0.35 -0.653 -0.838 0.124 0.972 -0.517 -0.203
oenopota_sp 0.33 -2.39 -0.254 -0.287 0.082 0.686 -0.456 0.0387 -0.66 -0.241 -0.0168 -0.38 -0.65 -0.0876 -0.159 0.397 0.0852 0.0713
oligochaeta 0.05 -2.96 -0.748 0.367 -0.35 0.364 -0.537 -0.973 0.178 0.641 -0.0921 -0.258 -0.455 -0.251 0.0128 0.338 0.535 -0.0576
ophelia_limacina 0.31 -2.61 0.35 -0.288 0.126 0.135 -0.275 -0.405 -0.387 0.298 -0.0124 -0.102 -0.649 -0.198 0.0453 0.0084 0.545 -0.11
opheliidae_spp 0.35 -3.64 0.459 -0.469 0.275 0.411 -0.353 -0.317 -0.486 0.0911 -0.0312 -0.25 -0.426 -0.484 -0.283 0.473 -0.131 0.0529
ophiopholis_aculeata 0.43 -3.74 0.222 -0.791 -0.12 0.552 -0.473 -0.559 -0.721 0.446 -0.387 0.161 -0.466 -0.564 0.122 0.384 -0.0543 0.194
ophiura_robusta 0.12 -6.08 -0.361 -2.27 0.0971 1.18 -1.15 -0.987 -2.28 0.501 -0.957 -0.55 -1.16 -1.85 0.371 1.68 -1.08 0.0443
orchomenella_minuta 0.33 -2.91 0.0905 -0.122 -0.074 0.258 -0.443 -0.346 -0.794 0.113 -0.00111 -0.17 -0.813 -0.0121 0.346 0.161 -0.219 -0.158
ostracoda 0 -1.28 0.378 -0.23 -0.53 0.0978 0.224 -0.659 -0.412 0.671 -0.377 0.529 -0.383 0.41 0.121 -0.559 0.618 0.0442
pagurus_sp 0.37 -3.84 -0.0169 -0.714 -0.119 0.374 -0.469 -0.871 -0.291 0.485 -0.242 0.236 -0.345 -0.605 0.00165 0.472 -0.188 -0.0972
pagurus_pubescens 0.55 -3.9 0.238 -1.08 0.111 0.163 -0.594 -0.721 -0.797 0.383 -0.27 0.0644 -0.569 -0.531 0.42 0.483 -0.614 -0.202
pandalus_montagui 0.16 -3.51 -0.139 -0.219 0.000365 0.517 -0.384 -0.416 -0.0741 0.182 0.358 -0.34 -0.0696 -0.57 -0.448 0.899 -0.0777 -0.0619
parathyasira_equalis 0.34 -3.79 0.0467 -0.582 -0.219 0.124 -0.412 -0.746 -0.303 0.579 -0.298 0.328 -0.315 -0.464 0.0167 0.283 -0.177 -0.0654
parvicardium_pinnulatum 0.48 -3.38 -0.0311 -1.03 -0.0972 0.498 -0.556 -0.756 -0.43 0.554 -0.393 0.115 -0.593 -0.669 0.161 0.692 -0.421 -0.175
periploma_leanum 0.29 -3.01 0.122 -0.332 -0.127 0.287 -0.38 -0.339 -0.295 0.405 -0.0145 0.0254 -0.591 -0.328 0.353 -0.033 0.561 -0.094
philine_lima 0.37 -3.89 -0.0111 -0.764 -0.128 0.391 -0.511 -0.832 -0.305 0.501 -0.264 0.262 -0.357 -0.621 0.0397 0.499 -0.188 -0.0953
philomedes_sp 0.39 -3.28 0.635 -0.515 0.0645 0.026 -0.21 -0.28 -0.35 0.118 -0.0268 -0.233 -0.0728 -0.106 -0.529 0.0796 0.194 0.103
pholoe_longa 0.36 -4.55 -0.723 -0.684 -0.124 0.532 -0.722 -1.09 -0.417 0.94 -0.384 -0.33 -0.813 -0.804 0.168 0.918 -0.405 -0.331
pholoe_sp 0.04 -0.0792 -0.337 0.419 -0.0287 0.149 0.187 -0.0277 -0.28 -0.108 -0.0638 0.0868 -0.337 0.178 -0.766 0.425 0.441 -0.0428
phoxocephalus_holbolli 0.16 -1.53 -0.365 0.0185 -0.0591 -0.248 -0.125 -0.246 0.163 -0.11 0.0452 -0.0637 -0.0855 0.17 0.138 -0.133 -0.723 -0.0586
polynoidae_spp 0.23 -1.33 0.15 0.277 -0.146 0.136 -0.474 -0.382 -0.507 -0.0725 0.148 -0.208 -0.774 -0.0608 0.42 -0.09 -0.131 -0.304
pontogeneia_inermis 0.37 -4.31 -0.612 -0.547 -0.0957 0.518 -0.625 -1.1 -0.391 0.948 -0.255 -0.312 -0.789 -0.824 -0.0225 1.05 -0.27 -0.305
pontoporeia_femorata 0.05 -1.54 -0.326 0.43 0.0523 0.74 -0.694 0.0978 0.298 -0.421 0.65 -0.616 -0.452 -0.0942 0.274 0.999 -0.55 -0.201
praxillella_praetermissa 0.16 -2.31 0.276 -0.277 -0.209 0.672 -0.362 -0.0354 0.228 0.205 -0.277 0.459 0.215 -0.374 0.216 0.557 0.569 0.0409
propebela_turricula 0.36 -2.78 0.815 -0.387 -0.197 0.194 -0.247 -0.232 -0.374 0.213 0.111 0.0334 -0.434 -0.137 0.158 -0.162 0.386 0.133
protomedeia_fasciata 0.2 -2.25 0.0466 0.314 0.141 0.219 -0.203 -0.483 0.878 -0.443 0.573 -0.518 0.302 -0.09 -1.24 0.775 -0.0191 0.282
protomedeia_grandimana 0.01 -0.319 0.639 0.612 -0.0172 -0.138 0.05 0.431 -0.341 -0.643 0.778 0.0784 -0.354 0.916 0.345 -0.467 -0.049 -0.0607
puncturella_noachina 0.45 -4.28 -0.287 -0.812 0.419 0.602 -0.56 -0.586 -0.834 0.349 -0.218 -0.596 -0.693 -1.05 -0.333 0.859 -0.323 -0.166
quasimelita_formosa 0.06 -1.16 0.628 -0.0972 -0.0964 0.34 -0.4 -0.188 -0.317 0.0689 -0.274 0.626 -0.23 0.502 0.86 -0.592 0.095 0.216
quasimelita_quadrispinosa 0.28 -3.73 -0.349 -0.183 -0.0927 0.38 -0.494 -0.748 -0.333 0.527 -0.0967 -0.254 -0.668 -0.477 -0.13 0.867 -0.145 -0.199
retusa_obtusa 0.33 -2.53 -0.609 -0.0448 -0.132 0.432 -0.304 0.123 0.508 -0.177 -0.0734 -0.319 -0.18 -0.274 -0.675 1.35 0.109 0.0587
sabellidae_spp 0.03 -3.17 -1.04 -0.481 -0.323 0.29 -0.281 -0.766 0.967 0.032 -0.123 -0.665 -0.518 -1.41 -0.0899 1.45 -0.034 -0.172
scoletoma_fragilis 0.4 -2.62 -0.923 0.267 0.0842 1.1 -0.0978 -0.223 -0.351 -0.046 -0.0323 -0.703 -0.78 -1.4 -0.465 2.19 -0.205 -0.305
scoletoma_sp 0.34 -3.61 -0.219 -0.715 -0.108 0.368 -0.455 -0.528 -0.145 0.218 -0.269 -0.256 0.26 -0.46 -0.294 0.565 -0.255 0.218
scoloplos_sp 0.36 -3.82 0.333 -0.681 -0.186 0.21 -0.418 -0.685 -0.406 0.557 -0.24 0.293 -0.32 -0.396 0.101 0.208 -0.181 0.0271
serripes_groenlandicus 0.18 -2.99 -0.072 0.145 0.273 0.254 -0.232 -0.273 -0.0275 0.194 0.166 -0.26 -0.354 -0.375 -0.357 0.4 0.437 -0.0989
sipuncula 0.23 -1.49 0.189 0.256 -0.16 0.468 0.187 0.329 -0.00901 -0.263 -0.0581 0.0211 -0.335 -0.00725 -0.24 0.477 0.302 -0.015
solamen_glandula 0.4 -4.39 -0.334 -0.724 0.0656 0.653 -0.586 -0.66 -0.766 0.413 -0.263 -0.465 -0.703 -0.992 -0.183 0.972 -0.386 -0.214
solariella_sp 0.51 -3.38 0.351 -0.831 0.154 0.685 -0.443 -0.27 -0.814 -0.18 -0.253 -0.0889 -0.516 -0.474 -0.208 0.827 0.0237 0.319
strongylocentrotus_sp 0.48 -4.52 -0.417 -1.15 0.211 0.965 -0.702 -0.543 -1.1 0.279 -0.436 -0.639 -0.678 -1.26 -0.291 1.09 -0.486 0.24
tachyrhynchus_erosus 0.35 -3.82 0.35 -0.654 -0.183 0.219 -0.424 -0.671 -0.389 0.527 -0.235 0.283 -0.303 -0.395 0.107 0.223 -0.191 0.0344
thracia_septentrionalis 0.11 -1.6 -0.839 -0.288 0.2 0.409 -0.0563 -0.396 -0.286 -0.146 -0.384 -0.0764 -0.269 -0.294 0.145 0.146 0.146 -0.0674
thyasira_gouldi 0.24 -1.27 0.882 0.0348 -0.128 0.126 -0.244 0.113 -1 0.0435 0.069 0.441 -0.763 0.629 0.459 -0.841 0.545 0.169
thyasira_sp 0.36 -4.2 0.518 -0.767 -0.279 0.142 -0.433 -0.788 -0.38 0.669 -0.281 0.483 -0.24 -0.388 0.161 0.133 -0.281 0.0599
trichotropis_bicarinata 0.22 -3.46 0.414 -0.413 -0.152 0.167 -0.301 -0.483 -0.179 0.277 -0.0739 -0.111 -0.0402 -0.0313 -0.281 0.182 0.0231 -0.0377
turritellopsis_stimpsoni 0.41 -3.39 0.36 -0.383 -0.0969 -0.118 -0.324 -0.549 -0.572 0.329 0.15 -0.108 -0.401 0.0105 -0.187 -0.184 -0.396 -0.0658
yoldia_myalis 0.23 -2.74 0.186 -0.206 0.0607 0.101 -0.337 -0.35 -0.369 0.253 -0.00821 0.134 -0.64 -0.406 0.162 0.319 0.132 -0.161
0.3 -3.03 -0.0685 -0.42 -0.0437 0.285 -0.395 -0.466 -0.363 0.24 -0.126 -0.124 -0.44 -0.346 -0.00564 0.381 -0.133 -0.0538
Ancillary information

Predictive power plot:

95 % confidence interval of the predictor coefficients estimated by the HMSC model:

Variance partitioning:

Trace plots:

Density plots:

Abiotic parameters
Model outcomes

Mean of the predictor coefficients estimated by the HMSC model:

Nakagawa’s pseudo-R2 and variables coefficients for each taxon
  Pseudo.R2 Intercept depth om gravel silt clay arsenic cadmium copper mercury
aceroides_aceroides_latipes 0.33 -1.58 0.0651 -0.501 -0.545 0.498 -0.98 -0.21 -0.0227 0.131 0.227
akanthophoreus_gracilis 0.08 -1.49 0.776 -0.719 -0.295 0.329 0.373 0.357 -0.447 -0.667 0.407
ameritella_agilis 0.09 -1.76 -0.0346 0.292 -0.0298 -0.335 0.161 -0.185 0.307 -0.25 -0.176
ameroculodes_edwardsi 0.52 -2.75 0.714 -0.62 0.156 0.409 -0.975 -0.335 -0.255 -0.295 0.0691
ampelisca_vadorum 0.6 -4.31 0.0297 -1.44 0.434 0.837 -1.6 -0.298 -0.6 0.0486 -0.306
amphipoda 0.03 -0.594 0.111 0.0602 0.0837 0.112 -0.255 -0.0639 0.369 -0.385 0.197
anonyx_lilljeborgi 0.42 -2.52 -0.645 -0.215 0.0526 0.336 -1.11 -0.337 -0.28 -0.0365 -0.139
anthozoa 0.59 -4.11 -0.108 -1.32 0.0418 1.18 -1.57 -0.34 -0.736 0.165 -0.428
arcteobia_anticostiensis 0.55 -3.81 -0.0755 -1.07 -0.0871 0.768 -1.43 -0.536 -0.254 -0.0459 -0.135
arrhoges_occidentalis 0.58 -4.08 0.195 -1.43 -0.173 0.647 -1.38 -0.344 -0.505 0.0622 -0.279
astarte_sp 0.63 -3.58 0.629 -1.39 -0.142 0.607 -1.28 -0.298 -0.708 -0.0437 -0.222
axinopsida_orbiculata 0.08 -1.41 0.578 0.107 -0.41 -0.342 0.0877 -0.061 0.425 -0.0873 -0.631
axiothella_catenata 0.44 -3 0.574 -0.722 -0.202 0.349 -1.03 -0.363 -0.0864 -0.185 0.687
bathymedon_longimanus 0.53 -3.44 -0.515 -0.341 -0.25 0.361 -1.2 -1.05 -0.0503 -0.0672 -0.445
bathymedon_obtusifrons 0.54 -4.19 -0.436 -1.84 -0.103 1.49 -1.97 -0.414 -0.45 0.985 -0.517
bipalponephtys_neotena 0 1.86 -0.0491 0.425 -0.201 0.273 -0.00959 -0.327 0.234 -0.285 0.322
boreochiton_ruber 0.69 -4.99 -0.564 -2.39 0.4 1.44 -2.2 -0.552 -1.28 0.0739 -0.668
brachydiastylis_sp 0.63 -4.02 0.519 -1.6 0.266 0.782 -1.5 -0.167 -0.648 -0.0767 -0.207
byblis_gaimardii 0.58 -3.45 0.649 -1.08 0.0131 0.264 -1.14 -0.269 -0.412 -0.362 0.0447
cancer_irroratus 0.55 -4.21 -0.746 -1.17 0.0555 0.691 -1.47 -0.479 -0.258 0.129 -0.313
caprella_septentrionalis 0.04 -5.46 -2.94 -0.625 -0.0634 -0.0235 -0.345 -1.17 -0.212 0.0926 -0.513
chaetodermatida 0.33 -1.43 0.669 -0.316 -0.457 -0.207 0.21 -0.324 -0.0268 -0.0687 0.173
chionoecetes_opilio 0.55 -3.4 0.593 -0.937 -0.055 0.857 -1.25 -0.3 -0.514 -0.0169 -0.161
chlamys_islandica 0.62 -4.55 -0.371 -1.67 -0.0412 1.04 -1.75 -0.474 -0.618 0.327 -0.452
chone_sp 0.52 -4.96 -1.23 -0.89 -0.207 0.777 -1.71 -1.11 -0.205 0.795 -0.509
ciliatocardium_ciliatum 0.48 -3.61 0.58 -0.933 -0.0398 0.195 -1.13 -0.248 -0.231 -0.0615 0.0227
cirripedia 0.45 -3.86 -0.568 -0.563 -0.184 0.264 -1.12 -0.738 0.46 -0.0204 -0.349
cistenides_granulata 0.42 -1.64 -0.0198 -0.466 0.119 0.41 -0.836 -0.237 -0.441 -0.456 -0.0215
cossura_longocirrata 0.4 -3.66 -0.227 -0.507 -0.121 0.83 -1.33 -0.102 0.149 0.303 -0.104
crassicorophium_bonellii 0.51 -3.45 0.244 -1.51 -0.182 0.365 -1.26 0.3 -0.428 0.195 -0.205
crenella_decussata 0.65 -3.26 0.0791 -1.53 0.372 0.732 -1.08 -0.18 -0.779 -0.308 -0.257
cumacea 0.2 -2.73 0.716 -0.466 -0.39 -0.00694 0.33 -0.0262 -0.0209 0.0885 0.0973
cyclocardia_borealis 0.64 -4.46 -0.0751 -1.57 0.112 1.46 -1.84 -0.367 -0.963 0.187 -0.545
cyrtodaria_siliqua 0.53 -3.89 0.218 -0.98 -0.23 0.421 -1.24 -0.554 -0.526 0.275 -0.302
diastylis_rathkei 0.25 -3.03 -0.963 -0.131 0.354 0.583 -1.49 -0.504 -0.665 0.387 -0.227
diastylis_sculpta 0.54 -3.64 -1.11 -0.512 -0.356 0.806 -1.67 -1.13 -0.269 0.972 -0.571
diastylis_sp 0.45 -3.38 0.606 -0.686 -0.116 0.682 -1.15 -0.365 -0.162 0.0641 -0.0795
echinarachnius_parma 0.62 -3.16 -0.34 -1.81 -0.255 0.728 -1.39 -0.76 -0.856 0.461 -0.962
edotia_montosa 0.46 -3.45 -1.06 -0.39 -0.268 -0.197 -1.03 -0.274 -0.114 0.378 -0.498
ennucula_tenuis 0.11 -0.224 0.907 -0.0545 -0.0106 0.174 0.229 0.0987 -0.354 -0.489 0.377
eteone_sp 0.57 -2.81 0.46 -1.53 0.206 0.736 -1.38 0.184 -0.417 0.151 -0.0129
euchone_sp 0.36 -5.58 -1.28 -1.16 -0.191 1.15 -2.03 -1.29 -0.298 1.13 -0.655
eudorella_emarginata 0.42 -2.3 -0.0816 0.0936 0.358 0.548 -0.987 -0.0788 0.0645 -0.0444 0.331
eudorellopsis_integra 0.01 0.639 0.455 0.427 -0.279 -0.14 -0.0504 0.778 0.0802 -0.814 1.05
euspira_pallida 0.57 -3.91 0.376 -1.32 -0.131 0.774 -1.44 -0.343 -0.55 0.176 -0.193
glycera_capitata 0.26 -3.15 -0.141 -1.11 0.386 0.675 0.184 -0.152 -0.333 0.0772 -0.207
glycera_sp 0.5 -2.99 -1.09 0.0105 0.0339 0.751 -1.23 -0.509 0.106 0.323 -0.475
goniada_maculata 0.34 -0.498 0.782 -0.0389 -0.526 0.0245 -0.627 -0.34 0.196 -0.157 0.315
guernea_prinassus_nordenskioldi 0.55 -4.38 -0.0596 -2.4 -0.102 1.69 -2.14 -0.13 -0.673 0.815 -0.567
halacaridae_spp 0.61 -4.42 -0.345 -1.61 -0.0415 1.01 -1.65 -0.447 -0.612 0.307 -0.417
haminoea_solitaria 0.41 -3.45 -0.0426 -0.533 -0.186 0.327 -1.07 -0.416 -0.156 -0.127 -0.131
hardametopa_carinata 0.51 -3.95 -0.689 -0.757 -0.137 0.707 -1.43 -0.787 -0.211 0.552 -0.38
harmothoe_sp 0.56 -3.08 0.483 -1.01 0.402 1.03 -1.37 -0.0925 -0.57 0.102 -0.194
harpacticoida 0.04 -0.239 -0.659 -0.247 -0.109 -0.174 -0.407 0.324 0.0835 -0.199 -0.156
hediste_diversicolor 0.47 -1.81 -1.5 0.339 0.0869 0.592 -0.262 0.0208 -0.2 -0.214 0.0198
heteranomia_squamula 0.67 -5.03 -0.517 -1.64 0.271 0.862 -1.81 -0.739 -0.818 0.162 -0.52
hiatella_arctica 0.64 -3.97 0.18 -1.34 -0.0158 0.203 -1.3 -0.641 -0.504 -0.0478 -0.203
holothuroidea 0.53 -3.44 -0.167 -0.611 -0.371 0.205 -1.14 -0.752 -0.63 0.316 -0.282
idotea_phosphorea 0.6 -4.09 -1.69 -0.998 0.247 1.08 -1.66 -0.528 -0.355 0.12 -0.491
ischyroceridae_spp 0.44 -3.58 -0.495 -0.287 -0.201 0.474 -1.22 -0.872 -0.171 0.528 -0.21
ischyrocerus_anguipes 0.08 -5.45 -2.58 0.469 -0.48 -0.384 -1.53 -1.56 0.516 0.513 -0.623
isopoda 0.49 -3.69 0.113 -0.793 -0.262 0.226 -1.16 -0.496 -0.298 0.00867 -0.169
lacuna_vincta 0.62 -4.67 -1.29 -1.18 0.056 0.526 -1.54 -0.743 -0.504 0.0412 -0.488
lamprops_fuscatus 0.42 -2.18 -0.136 -0.686 -0.376 0.265 -0.876 -0.557 -0.284 0.401 -0.22
lamprops_quadriplicata 0.44 -3.51 -0.727 -0.804 -0.163 -0.0341 -1.13 0.0159 -0.0812 0.0532 -0.159
lepeta_caeca 0.67 -3.35 0.0429 -1.64 0.521 1.12 -1.66 -0.215 -0.695 -0.0454 -0.317
leucon_leucon_nasicoides 0.01 -0.457 1.06 0.692 -0.261 -0.364 0.0551 0.151 0.0811 -0.32 0.751
littorina_littorea 0.45 -3.09 -1.15 0.0758 -0.0646 0.531 -1.12 -0.66 -0.0754 0.279 -0.487
lumbrineridae_spp 0.38 -3.2 0.458 -0.299 -0.261 0.252 -0.995 -0.672 0.767 -0.34 -0.25
lysianassidae_spp 0.54 -3.99 -0.688 -0.9 -0.165 0.778 -1.08 -0.808 -0.692 0.483 -0.711
macoma_calcarea 0 0.995 -0.24 0.103 -0.154 0.0442 -0.243 -0.102 -0.161 -0.195 -0.222
maera_danae 0.52 -4.1 -0.816 -0.797 -0.125 0.711 -1.44 -0.784 -0.205 0.513 -0.373
maldane_sarsi 0.38 -2.85 0.502 -0.394 -0.129 0.749 -0.997 0.103 -0.195 0.336 -0.157
maldanidae_spp 0.11 -1.73 0.921 -0.128 -0.711 0.69 0.181 -0.0854 -0.0982 0.559 0.255
monoculopsis_longicornis 0.57 -3.77 0.351 -1.07 -0.386 0.351 -1.3 -0.5 -0.731 0.0811 -0.0647
muculus_musculus_discors 0.61 -4.36 -0.34 -1.59 -0.0525 1 -1.69 -0.431 -0.6 0.298 -0.409
mytilus_sp 0.02 -2.48 -0.211 0.103 -0.226 0.0978 -1.03 -1.43 -0.0718 0.132 -0.296
nematoda 0.01 -1.05 0.63 -2.16 0.0247 0.392 -1.19 0.793 -0.556 -0.908 0.137
nemertea 0.5 -3.65 0.281 -1.44 0.579 0.628 -1.51 -0.133 -0.344 -0.239 0.685
neoleanira_tetragona 0.48 -3.96 -0.626 -0.698 -0.219 0.0797 -1.16 -0.523 -0.049 -0.11 -0.159
nephtyidae_spp 0.29 -2.03 0.323 -0.286 -0.487 -0.118 -0.757 0.239 0.0571 -0.352 0.112
nephtys_caeca 0.36 -2.22 -0.639 0.0124 0.184 -0.133 -0.855 -0.21 0.204 -0.0805 -0.407
nephtys_incisa 0.19 -0.964 0.752 0.141 -0.0668 0.0368 0.196 -0.188 -0.186 -0.126 0.419
nephtys_sp 0.43 -3 0.46 -0.867 -0.221 0.6 -1.15 -0.13 -0.171 0.161 0.00514
nuculana_minuta 0.51 -2.02 0.534 -0.712 0.135 0.722 -1.1 -0.155 -0.256 -0.0181 0.0872
nymphonidae_spp 0.64 -4.76 -0.412 -1.5 0.17 0.811 -1.73 -0.656 -0.69 0.178 -0.463
oenopota_sp 0.57 -2.48 -0.199 -0.889 0.0867 1.03 -1.36 -0.0445 -0.692 0.0244 -0.18
oligochaeta 0.09 -3.22 -0.76 0.162 -0.534 0.525 -1.5 -0.868 0.196 0.703 0.169
ophelia_limacina 0.39 -2.39 0.4 -0.633 0.138 0.361 -1.03 -0.15 -0.102 0.153 0.194
opheliidae_spp 0.55 -3.46 0.645 -1.02 0.404 0.814 -1.29 -0.155 -0.392 -0.108 -0.147
ophiopholis_aculeata 0.61 -3.74 0.16 -1.36 -0.0412 1.1 -1.52 -0.328 -0.683 0.0788 -0.34
ophiura_robusta 0.18 -6.9 -0.188 -3.83 0.283 2.06 -3.07 -0.567 -2.19 0.139 -1.17
orchomenella_minuta 0.4 -2.74 -0.0223 -0.0544 -0.196 0.303 -0.942 -0.576 -0.39 0.056 -0.0816
ostracoda 0 -1.25 0.492 -0.572 -0.731 0.321 0.223 -0.433 -0.26 0.316 0.0266
pagurus_sp 0.55 -3.85 -0.0762 -1.1 -0.0622 0.787 -1.41 -0.53 -0.268 -0.0959 -0.127
pagurus_pubescens 0.62 -3.78 0.323 -1.34 0.117 0.265 -1.28 -0.485 -0.439 -0.116 -0.125
pandalus_montagui 0.43 -3.34 -0.181 -0.57 -0.0837 0.784 -1.26 -0.296 -0.304 0.29 0.27
parathyasira_equalis 0.43 -3.61 -0.0892 -0.555 -0.227 0.306 -1.12 -0.458 -0.138 -0.14 -0.134
parvicardium_pinnulatum 0.52 -3.07 0.0874 -1.37 -0.0608 0.816 -1.03 -0.417 -0.251 0.236 -0.308
periploma_leanum 0.4 -2.82 0.19 -0.699 -0.221 0.617 -1.13 0.0964 0.00321 0.125 0.306
philine_lima 0.54 -3.91 -0.0774 -1.12 -0.0603 0.78 -1.38 -0.541 -0.257 -0.0824 -0.104
philomedes_sp 0.62 -3.51 0.899 -1.38 0.127 0.434 -1.24 -0.117 -0.505 -0.154 -0.0798
pholoe_longa 0.52 -4.96 -1.07 -0.976 -0.174 0.962 -1.84 -1.12 -0.252 0.928 -0.526
pholoe_sp 0.02 -0.0493 -0.212 0.284 -0.0533 0.13 0.12 -0.242 -0.444 0.0668 -0.158
phoxocephalus_holbolli 0.11 -1.47 -0.59 0.156 -0.0733 -0.292 -0.0428 -0.41 0.317 -0.219 -0.161
polynoidae_spp 0.27 -1.32 0.0891 0.167 -0.243 0.273 -1 -0.457 -0.0881 -0.0604 0.143
pontogeneia_inermis 0.52 -4.24 -0.741 -0.781 -0.186 0.865 -1.54 -1.08 -0.236 0.908 -0.334
pontoporeia_femorata 0.05 -1.62 -0.584 0.435 -0.0131 0.785 -1.18 -0.321 0.132 0.423 0.223
praxillella_praetermissa 0.26 -2.59 0.315 -0.55 -0.167 1.04 -1.32 0.293 -0.0169 0.338 -0.0848
propebela_turricula 0.48 -2.64 0.923 -0.897 -0.272 0.541 -1.03 0.0173 -0.199 0.0638 0.268
protomedeia_fasciata 0.27 -2.41 0.144 0.326 0.18 0.0925 -1.06 -0.893 0.534 -0.114 0.287
protomedeia_grandimana 0.01 -0.307 0.475 0.497 -0.164 -0.179 0.0662 0.0565 -0.241 -0.195 0.649
puncturella_noachina 0.65 -4.1 0.0365 -1.59 0.572 1.27 -1.77 -0.2 -0.857 0.0282 -0.392
quasimelita_formosa 0.09 -1.32 0.576 -0.254 -0.0645 0.628 -0.931 -0.243 -0.0896 0.0482 -0.144
quasimelita_quadrispinosa 0.42 -3.63 -0.535 -0.199 -0.193 0.467 -1.18 -0.897 -0.195 0.554 -0.231
retusa_obtusa 0.39 -2.53 -0.81 -0.157 -0.312 0.294 -0.793 0.00591 0.264 0.281 -0.44
sabellidae_spp 0.02 -3.3 -0.934 -0.561 -0.649 0.238 -0.109 -0.617 1.12 0.0219 -0.0642
scoletoma_fragilis 0.31 -2.38 -0.899 0.353 -0.0951 1.03 0.0279 -0.404 -0.457 0.347 -0.393
scoletoma_sp 0.59 -3.9 -0.359 -1.45 -0.122 0.875 -1.47 -0.363 -0.575 0.249 -0.442
scoloplos_sp 0.46 -3.53 0.326 -0.845 -0.199 0.464 -1.14 -0.368 -0.199 -0.0621 -0.0844
serripes_groenlandicus 0.34 -2.83 -0.165 -0.0154 0.373 0.437 -1.01 -0.186 0.0378 0.0587 0.16
sipuncula 0.24 -1.44 0.255 0.0573 -0.316 0.504 0.188 0.238 -0.0557 -0.0173 -0.112
solamen_glandula 0.6 -4.15 -0.119 -1.34 0.0505 1.17 -1.6 -0.383 -0.72 0.177 -0.442
solariella_sp 0.69 -3.85 0.549 -1.77 0.302 1.15 -1.79 -0.348 -0.882 -0.161 -0.399
strongylocentrotus_sp 0.67 -4.98 -0.213 -2.63 0.507 1.98 -2.36 -0.137 -1.41 0.206 -0.752
tachyrhynchus_erosus 0.48 -3.51 0.351 -0.888 -0.178 0.513 -1.19 -0.327 -0.216 -0.0525 -0.0933
thracia_septentrionalis 0.1 -1.59 -0.941 -0.271 0.263 0.473 0.00293 -0.232 -0.233 -0.297 -0.206
thyasira_gouldi 0.2 -1.26 0.955 -0.444 -0.148 0.49 -0.934 0.137 -0.491 -0.222 0.362
thyasira_sp 0.46 -4.32 0.537 -1.04 -0.24 0.574 -1.38 -0.468 -0.197 -0.13 -0.0834
trichotropis_bicarinata 0.5 -3.66 0.492 -0.994 -0.194 0.565 -1.26 -0.402 -0.34 0.165 -0.148
turritellopsis_stimpsoni 0.5 -3.11 0.512 -0.771 -0.126 0.172 -1.03 -0.386 -0.364 -0.103 0.176
yoldia_myalis 0.31 -2.58 0.226 -0.262 0.0573 0.156 -0.923 -0.146 -0.0664 -0.0678 0.118
0.4 -3.01 -0.0897 -0.736 -0.07 0.539 -1.06 -0.353 -0.276 0.0838 -0.149
Ancillary information

Predictive power plot:

95 % confidence interval of the predictor coefficients estimated by the HMSC model:

Variance partitioning:

Trace plots:

Density plots:

Exposure indices
Model outcomes

Mean of the predictor coefficients estimated by the HMSC model:

Nakagawa’s pseudo-R2 and variables coefficients for each taxon
  Pseudo.R2 Intercept depth aquaculture dredging runoff sewers structures shipping fisheries
aceroides_aceroides_latipes 0.14 -1.46 0.0163 0.342 -0.755 0.131 0.0588 0.0412 0.478 0.0398
akanthophoreus_gracilis 0.09 -1.41 0.523 0.611 -0.266 0.913 -0.184 -0.968 0.582 0.477
ameritella_agilis 0.12 -1.85 0.0185 -0.0242 -0.0221 0.0924 0.0526 -0.355 -0.427 0.0364
ameroculodes_edwardsi 0.3 -2.48 0.515 0.13 -0.221 0.422 -0.697 0.0679 -0.135 -0.00695
ampelisca_vadorum 0.27 -4.01 -0.203 -0.648 -1.09 -1.41 -1.28 2.11 -0.00919 0.13
amphipoda 0.03 -0.592 0.128 0.169 0.278 0.508 0.279 -1.04 -0.0478 0.0277
anonyx_lilljeborgi 0.24 -2.37 -0.581 -0.0271 -0.993 -0.75 -0.665 1.16 0.108 -0.318
anthozoa 0.31 -4.29 -0.575 -0.898 -1.27 -1.55 -1.42 2.37 -0.31 -0.17
arcteobia_anticostiensis 0.27 -3.74 -0.248 0.277 -1.04 -0.568 -0.78 1.48 -0.168 0.00917
arrhoges_occidentalis 0.22 -3.74 -0.251 -0.604 -1.01 -0.953 -1.28 1.58 0.189 -0.0874
astarte_sp 0.38 -3.09 0.271 -0.588 -1.19 0.122 -1.13 0.795 -0.213 -0.0149
axinopsida_orbiculata 0.06 -1.45 0.456 0.0042 0.479 0.36 -0.281 -0.255 -0.147 0.295
axiothella_catenata 0.21 -3.16 0.3 -0.179 -0.711 -0.782 -0.175 0.598 0.468 0.0333
bathymedon_longimanus 0.32 -3.04 -0.602 0.5 -0.251 -0.364 -0.872 1.45 -0.225 -0.0224
bathymedon_obtusifrons 0.3 -4.08 -0.402 -0.408 -1.75 -1.08 0.473 1.83 -0.535 -0.368
bipalponephtys_neotena 0 1.87 0.0379 0.532 -0.203 0.273 0.38 -0.24 0.39 -0.0179
boreochiton_ruber 0.45 -4.67 -0.948 -0.725 -1.51 -2.18 -2.37 4.04 -0.668 0.329
brachydiastylis_sp 0.3 -3.57 0.0094 -0.541 -1.01 -1 -1.41 1.69 0.304 0.193
byblis_gaimardii 0.45 -3.59 0.124 -0.257 -1.04 -0.402 -1.69 1.34 0.0751 0.343
cancer_irroratus 0.27 -3.69 -0.615 -0.4 -0.858 0.0697 -0.0596 0.364 -0.515 -0.204
caprella_septentrionalis 0.05 -6.81 -2.73 -1.34 -2.14 -0.139 -0.797 1.72 -1.66 -0.648
chaetodermatida 0.2 -1.31 0.66 0.218 -0.161 0.76 0.216 -0.554 0.00149 0.194
chionoecetes_opilio 0.27 -3.12 0.406 0.0387 -0.977 -0.433 -0.755 1.08 0.118 -0.0939
chlamys_islandica 0.3 -4.22 -0.489 -0.573 -1.11 -1.12 -1.09 1.91 -0.254 0.33
chone_sp 0.41 -5.2 -1.07 -0.547 -1.81 -0.155 -0.784 1.66 -0.847 0.0454
ciliatocardium_ciliatum 0.26 -3.77 0.199 -0.575 -1.09 -1.24 -0.917 1.81 -0.0841 0.087
cirripedia 0.38 -4.78 -1.19 -0.95 -1.46 -1.9 -0.953 2.89 -0.551 -0.226
cistenides_granulata 0.2 -1.56 -0.117 -0.0651 -0.52 -0.713 -0.981 1.39 0.0422 0.169
cossura_longocirrata 0.21 -3.6 -0.421 -0.465 -0.786 -0.63 -0.507 0.597 0.761 -0.145
crassicorophium_bonellii 0.19 -3.35 -0.345 -0.695 -0.785 -1.36 -1.17 1.95 0.268 -0.000464
crenella_decussata 0.45 -2.75 -0.188 -0.155 -1.23 -1.5 -1.53 2.71 -0.0901 0.0614
cumacea 0.18 -2.84 0.513 -0.283 -0.348 -0.99 0.435 0.901 -0.112 -0.199
cyclocardia_borealis 0.37 -4.58 -0.654 -1.11 -1.44 -1.92 -1.8 2.85 -0.4 -0.228
cyrtodaria_siliqua 0.14 -3.22 0.0436 -0.215 -0.124 -0.458 -0.512 1.22 -0.17 -0.0442
diastylis_rathkei 0.16 -3.1 -1.1 -1.08 0.19 -0.758 -1.63 1.39 -0.00223 -0.181
diastylis_sculpta 0.4 -3.86 -1.28 -0.908 -1.66 -1.2 -1.78 2.92 -0.0482 -0.335
diastylis_sp 0.26 -3.58 0.417 -0.233 -1.06 -0.824 -0.656 1.49 -0.207 0.0165
echinarachnius_parma 0.24 -2.27 -0.523 0.0948 -0.627 -0.708 -1.37 2.19 -0.109 0.0721
edotia_montosa 0.45 -3.93 -1.31 -0.848 -0.663 -0.987 -0.708 1.87 -0.839 -0.419
ennucula_tenuis 0.12 -0.234 0.928 0.524 -0.00332 0.522 -0.132 -0.341 0.531 0.262
eteone_sp 0.27 -2.47 0.166 -0.525 -0.672 -1.07 -0.126 0.75 0.271 0.197
euchone_sp 0.21 -5.44 -1.01 -0.586 -2.12 -0.381 -0.665 1.93 -0.531 -0.332
eudorella_emarginata 0.31 -2.44 -0.176 -0.455 -0.168 -1.15 0.222 0.68 0.431 -0.275
eudorellopsis_integra 0.01 0.64 0.381 0.117 0.323 0.135 0.136 -0.756 1.23 -0.127
euspira_pallida 0.2 -3.33 0.0833 -0.209 -0.838 0.464 -0.696 0.136 0.102 -0.0174
glycera_capitata 0.24 -3.38 -0.482 -0.745 -1.07 -1.58 -0.573 1.8 -0.0973 -0.0267
glycera_sp 0.41 -3.41 -1.05 -1.01 -0.957 -0.784 -0.477 0.871 -0.595 -0.5
goniada_maculata 0.21 -0.406 0.759 0.229 -0.25 0.115 0.156 -0.0416 0.27 0.123
guernea_prinassus_nordenskioldi 0.2 -4.16 -0.244 -0.717 -1.58 -1.83 0.0326 2.49 -0.603 -0.232
halacaridae_spp 0.28 -4.16 -0.495 -0.54 -1.09 -1.08 -1.05 1.84 -0.219 0.329
haminoea_solitaria 0.28 -3.78 -0.177 0.259 -1.05 -0.563 -0.802 1.47 -0.196 0.0255
hardametopa_carinata 0.33 -4.07 -0.726 -0.485 -1.37 -0.422 -0.583 1.46 -0.389 -0.197
harmothoe_sp 0.27 -3 0.279 -0.618 -0.925 -1.48 -0.621 1.65 0.141 0.0983
harpacticoida 0.03 -0.278 -0.525 -0.278 -0.367 0.605 -0.0809 -0.382 0.0989 -0.0719
hediste_diversicolor 0.49 -2.3 -1.09 -0.73 0.146 -0.863 0.366 0.849 -0.68 -0.554
heteranomia_squamula 0.34 -4.52 -0.622 -0.597 -1.35 -1.28 -1.26 2.9 -0.413 -0.119
hiatella_arctica 0.35 -3.36 -0.0207 -0.476 -1.49 -1.22 -0.797 2.65 -0.465 -0.176
holothuroidea 0.4 -3.4 -0.448 -0.336 -0.779 0.816 -1.38 0.494 -0.242 0.0825
idotea_phosphorea 0.36 -3.85 -0.811 -0.375 -0.823 0.86 0.106 -0.415 -0.813 -0.266
ischyroceridae_spp 0.26 -3.86 -0.405 -0.33 -1.31 -0.514 -0.598 1.66 -0.263 -0.196
ischyrocerus_anguipes 0.12 -6.87 -2.79 -1.82 -2.34 -1.24 -1.46 2.82 -1.91 -0.74
isopoda 0.29 -3.77 -0.206 -0.309 -0.899 0.2 -1.02 0.627 -0.143 0.269
lacuna_vincta 0.44 -4.51 -1.36 -0.794 -1.35 -0.614 -0.721 1.67 -0.748 -0.272
lamprops_fuscatus 0.11 -1.85 -0.157 0.399 -0.223 0.374 -0.206 0.367 0.215 0.225
lamprops_quadriplicata 0.35 -3.64 -0.964 -0.622 -0.698 -0.663 0.00778 0.967 -0.518 -0.347
lepeta_caeca 0.45 -3.11 -0.263 -0.584 -1.26 -0.849 -1.04 1.6 -0.784 0.101
leucon_leucon_nasicoides 0.01 -0.486 1.07 0.224 -0.0503 -0.983 0.712 0.549 0.421 0.0301
littorina_littorea 0.39 -3.53 -0.511 -0.449 -1.07 0.418 -0.013 0.363 -1.06 -0.299
lumbrineridae_spp 0.32 -4.4 0.0815 -0.635 -1.65 -2.2 -0.823 3.2 -0.715 -0.113
lysianassidae_spp 0.4 -3.88 -0.585 0.127 -0.977 -0.754 -1.38 2.38 -0.332 0.0465
macoma_calcarea 0 1.03 0.0388 0.255 0.144 1.15 0.0382 -0.904 -0.0818 0.0961
maera_danae 0.35 -4.08 -0.717 -0.453 -1.33 -0.314 -0.619 1.37 -0.541 -0.0265
maldane_sarsi 0.19 -3 0.466 -0.197 -0.738 -1.09 0.384 1.27 -0.0708 0.0793
maldanidae_spp 0.1 -1.54 0.908 0.582 -0.676 -0.085 0.702 0.211 0.83 -0.143
monoculopsis_longicornis 0.35 -3.08 0.00941 -0.138 -0.68 1.41 -0.901 -0.673 -0.0165 0.204
muculus_musculus_discors 0.28 -4.18 -0.51 -0.575 -1.03 -1.12 -1.04 1.88 -0.268 0.329
mytilus_sp 0.02 -2.64 -0.00288 -0.0498 -0.841 -0.115 -0.516 1.7 -1.16 -0.168
nematoda 0 -0.728 0.155 0.14 -1.24 0.612 -0.598 -0.559 -0.0636 0.289
nemertea 0.24 -3.85 -0.195 -0.71 -0.928 -1.52 -0.786 1.3 0.842 0.149
neoleanira_tetragona 0.27 -3.68 -0.631 -0.421 -0.833 0.0301 -0.0453 0.33 -0.519 -0.208
nephtyidae_spp 0.23 -1.93 0.183 0.06 0.536 0.591 -0.0204 -0.923 -0.43 0.313
nephtys_caeca 0.28 -2.37 -0.855 -0.462 -0.193 -0.581 -0.986 1.1 -0.196 -0.0509
nephtys_incisa 0.2 -0.955 0.806 0.283 -0.165 0.292 -0.00922 -0.17 0.441 -0.0943
nephtys_sp 0.2 -2.86 0.348 -0.271 -0.785 -0.249 -0.175 0.315 0.413 -0.149
nuculana_minuta 0.34 -1.96 0.412 0.104 -1.08 -0.771 -0.0693 1.08 0.283 0.2
nymphonidae_spp 0.28 -4.03 -0.527 -0.492 -1.16 -0.977 -1.02 2.25 -0.305 -0.129
oenopota_sp 0.19 -2.32 -0.331 -0.569 -0.645 -0.0864 -0.698 0.413 0.12 0.000351
oligochaeta 0.05 -3.34 -0.892 -0.5 -0.446 0.00364 0.312 0.163 0.561 -0.168
ophelia_limacina 0.27 -2.45 0.325 -0.324 -0.944 -0.496 -0.388 0.721 0.633 -0.0305
opheliidae_spp 0.32 -3.57 0.336 -0.462 -1.03 -1.32 -1.09 1.93 -0.00731 0.143
ophiopholis_aculeata 0.3 -3.41 0.0655 0.085 -0.87 -0.75 -0.833 1.4 0.0415 0.354
ophiura_robusta 0.07 -6.45 -0.91 -1.35 -2.73 -3.95 -3.56 6.73 -0.865 0.378
orchomenella_minuta 0.23 -2.76 0.15 -0.259 -1.08 0.171 -0.182 0.559 -0.219 -0.19
ostracoda 0 -1.18 0.395 0.492 -0.597 0.883 -0.216 -0.323 0.537 0.0963
pagurus_sp 0.26 -3.65 -0.222 0.264 -1.02 -0.552 -0.765 1.41 -0.134 0.0162
pagurus_pubescens 0.38 -3.28 0.0779 -0.0677 -1.36 -1.03 -0.865 2.41 -0.491 -0.114
pandalus_montagui 0.14 -3.17 -0.379 -0.429 0.186 -1 -0.29 1.03 0.12 -0.119
parathyasira_equalis 0.3 -3.83 -0.195 0.313 -1.09 -0.544 -0.824 1.5 -0.214 0.0474
parvicardium_pinnulatum 0.41 -3.12 -0.157 0.00925 -1.51 -0.707 -0.807 1.86 -0.476 -0.0918
periploma_leanum 0.26 -2.92 0.0448 -0.117 -0.628 -0.271 0.354 0.0475 0.597 -0.119
philine_lima 0.26 -3.63 -0.21 0.282 -1.01 -0.493 -0.709 1.34 -0.141 0.0169
philomedes_sp 0.27 -2.75 0.411 -0.496 -0.249 -0.571 -1.12 0.966 0.408 0.153
pholoe_longa 0.35 -4.97 -0.94 -0.534 -1.87 -0.407 -0.647 1.81 -0.529 -0.284
pholoe_sp 0.02 -0.0385 -0.157 0.0522 -0.237 -0.0631 -0.799 0.655 0.345 -0.15
phoxocephalus_holbolli 0.16 -1.53 -0.411 -0.0455 -0.232 0.22 0.0129 0.027 -0.859 -0.0357
polynoidae_spp 0.15 -1.25 0.365 -0.255 -0.882 -0.0585 0.14 0.17 -0.14 -0.402
pontogeneia_inermis 0.38 -4.63 -0.783 -0.474 -1.85 -0.51 -0.78 1.99 -0.368 -0.28
pontoporeia_femorata 0.05 -1.77 -0.398 -0.86 0.000362 -1.09 1.19 0.809 -0.547 -0.605
praxillella_praetermissa 0.14 -2.27 0.307 0.542 0.286 -0.709 0.614 0.581 0.491 -0.144
propebela_turricula 0.32 -2.5 0.699 -0.181 -0.66 -0.472 -0.121 0.381 0.456 0.143
protomedeia_fasciata 0.07 -2.42 -0.105 -0.396 0.39 -0.651 -0.551 0.579 -0.0728 0.149
protomedeia_grandimana 0.01 -0.319 0.782 0.175 0.136 0.514 0.781 -0.739 -0.00988 -0.191
puncturella_noachina 0.33 -3.92 -0.373 -0.971 -1.17 -1.87 -1.64 2.53 -0.0327 0.0049
quasimelita_formosa 0.04 -1.12 0.804 0.763 -0.195 0.964 0.82 -0.829 0.0375 0.29
quasimelita_quadrispinosa 0.27 -3.89 -0.413 -0.323 -1.35 -0.416 -0.59 1.63 -0.266 -0.194
retusa_obtusa 0.33 -2.67 -0.87 -0.424 -0.179 -0.946 -0.397 1.68 -0.0663 -0.106
sabellidae_spp 0.04 -4.34 -1.89 -1.47 -1.61 -2.47 -0.245 2.51 -0.127 -0.32
scoletoma_fragilis 0.39 -2.73 -0.687 -1.12 -0.629 -2.74 -0.345 3.1 -0.541 -0.669
scoletoma_sp 0.19 -3.09 -0.515 -0.451 0.261 -0.53 -0.792 0.842 -0.125 0.269
scoloplos_sp 0.28 -3.64 0.106 0.204 -1.01 -0.58 -0.676 1.41 -0.197 0.114
serripes_groenlandicus 0.19 -2.8 -0.13 -0.291 -0.105 -0.936 -0.159 0.818 0.617 -0.0994
sipuncula 0.15 -1.44 0.353 0.0188 -0.0758 -0.625 0.306 0.5 0.0195 -0.317
solamen_glandula 0.3 -4.21 -0.522 -0.869 -1.25 -1.56 -1.38 2.37 -0.316 -0.152
solariella_sp 0.42 -3.33 0.157 -0.243 -1.17 -1.25 -1.68 2.63 0.276 0.462
strongylocentrotus_sp 0.42 -4.64 -0.848 -1.25 -1.41 -2.35 -2.2 3.27 -0.388 0.415
tachyrhynchus_erosus 0.27 -3.64 0.103 0.219 -0.978 -0.554 -0.659 1.36 -0.179 0.137
thracia_septentrionalis 0.07 -1.63 -0.911 -0.0813 -0.272 -0.0439 -0.475 0.244 0.095 0.0538
thyasira_gouldi 0.18 -1.23 1 0.426 -0.86 0.815 0.0331 -0.641 0.651 0.223
thyasira_sp 0.3 -4.25 0.206 0.432 -1.16 -0.6 -0.731 1.58 -0.257 0.235
trichotropis_bicarinata 0.08 -3.05 0.257 -0.236 -0.147 0.00153 -0.571 0.457 0.0667 -0.0545
turritellopsis_stimpsoni 0.35 -3.11 0.175 -0.226 -0.941 0.0221 -0.92 0.672 -0.341 0.0312
yoldia_myalis 0.23 -2.76 0.192 0.12 -1.13 -0.927 -0.265 1.42 0.227 -0.129
0.24 -2.97 -0.218 -0.288 -0.807 -0.564 -0.571 1.15 -0.119 -0.0343
Ancillary information

Predictive power plot:

95 % confidence interval of the predictor coefficients estimated by the HMSC model:

Variance partitioning:

Trace plots:

Density plots:

3. Predictions of species richness

3.1. Generalized Linear Models

All variables

Prediction of specific richness based on this model:

Difference between predicted and observed taxa richness
station_id observed predicted difference
127 1 11 10
128 12 4 -8
129 15 3 -12
130 4 4 0
131 10 2 -8
132 10 16 6
134 9 10 1
135 8 13 5
136 9 6 -3
137 11 6 -5
138 12 8 -4
139 13 10 -3
140 14 9 -5
141 14 12 -2
142 22 6 -16
143 12 7 -5
144 12 8 -4
145 11 5 -6
146 10 7 -3
147 9 7 -2
148 20 13 -7
149 17 15 -2
150 11 15 4
151 11 19 8
152 15 19 4
153 17 18 1
154 11 21 10
155 20 21 1
156 22 17 -5
157 5 13 8
158 19 10 -9
159 17 11 -6
160 14 7 -7
161 8 3 -5
162 11 6 -5
163 13 7 -6
164 14 4 -10
165 5 6 1
166 7 6 -1
167 6 3 -3
168 19 6 -13
169 10 17 7
170 22 4 -18
171 14 11 -3
172 10 14 4
173 18 25 7
174 19 19 0
175 13 15 2
176 17 25 8
177 9 16 7
178 19 14 -5
179 17 18 1
180 12 19 7
181 21 23 2
182 18 14 -4
183 6 16 10
184 12 14 2
185 21 10 -11
186 14 14 0
187 7 7 0
188 5 32 27
189 16 16 0
190 20 8 -12
191 9 18 9
192 12 9 -3
193 2 7 5
194 7 NA NA
195 9 17 8
196 5 6 1
197 13 5 -8
198 14 15 1
199 7 16 9
200 15 10 -5
201 12 14 2
202 20 4 -16
203 19 10 -9
204 17 20 3
205 12 4 -8
206 11 7 -4
207 17 11 -6
208 18 12 -6
209 5 9 4
211 23 23 0
212 4 9 5
214 24 14 -10
215 6 13 7
216 6 13 7
217 12 18 6
218 17 12 -5
219 9 13 4
220 11 14 3
221 16 18 2
222 17 17 0
223 17 17 0
224 11 21 10
225 21 17 -4
226 17 15 -2
228 6 8 2
229 17 9 -8
230 17 13 -4
231 9 9 0
235 19 20 1
236 7 19 12
237 14 4 -10
238 7 3 -4
239 14 19 5
240 9 15 6
241 12 18 6
Abiotic parameters

Prediction of specific richness based on this model:

Difference between predicted and observed taxa richness
station_id observed predicted difference
127 1 3 2
128 12 2 -10
129 15 4 -11
130 4 5 1
131 10 4 -6
132 10 6 -4
134 9 6 -3
135 8 7 -1
136 9 7 -2
137 11 8 -3
138 12 8 -4
139 13 7 -6
140 14 8 -6
141 14 7 -7
142 22 6 -16
143 12 6 -6
144 12 6 -6
145 11 6 -5
146 10 6 -4
147 9 6 -3
148 20 6 -14
149 17 6 -11
150 11 4 -7
151 11 4 -7
152 15 4 -11
153 17 4 -13
154 11 4 -7
155 20 4 -16
156 22 4 -18
157 5 4 -1
158 19 4 -15
159 17 3 -14
160 14 7 -7
161 8 3 -5
162 11 3 -8
163 13 4 -9
164 14 4 -10
165 5 8 3
166 7 5 -2
167 6 5 -1
168 19 4 -15
169 10 14 4
170 22 4 -18
171 14 8 -6
172 10 8 -2
173 18 10 -8
174 19 13 -6
175 13 14 1
176 17 9 -8
177 9 14 5
178 19 15 -4
179 17 8 -9
180 12 15 3
181 21 9 -12
182 18 10 -8
183 6 11 5
184 12 12 0
185 21 10 -11
186 14 12 -2
187 7 8 1
188 5 53 48
189 16 13 -3
190 20 3 -17
191 9 3 -6
192 12 4 -8
193 2 3 1
194 7 NA NA
195 9 7 -2
196 5 8 3
197 13 3 -10
198 14 13 -1
199 7 4 -3
200 15 9 -6
201 12 8 -4
202 20 5 -15
203 19 12 -7
204 17 5 -12
205 12 6 -6
206 11 6 -5
207 17 10 -7
208 18 10 -8
209 5 4 -1
211 23 6 -17
212 4 7 3
214 24 9 -15
215 6 8 2
216 6 9 3
217 12 10 -2
218 17 8 -9
219 9 8 -1
220 11 8 -3
221 16 12 -4
222 17 12 -5
223 17 13 -4
224 11 8 -3
225 21 11 -10
226 17 11 -6
228 6 8 2
229 17 6 -11
230 17 5 -12
231 9 8 -1
235 19 12 -7
236 7 4 -3
237 14 5 -9
238 7 3 -4
239 14 4 -10
240 9 4 -5
241 12 3 -9
Exposure indices

Prediction of specific richness based on this model:

Difference between predicted and observed taxa richness
station_id observed predicted difference
127 1 31 30
128 12 9 -3
129 15 8 -7
130 4 8 4
131 10 7 -3
132 10 16 6
134 9 19 10
135 8 10 2
136 9 5 -4
137 11 6 -5
138 12 10 -2
139 13 7 -6
140 14 8 -6
141 14 7 -7
142 22 3 -19
143 12 3 -9
144 12 4 -8
145 11 3 -8
146 10 3 -7
147 9 4 -5
148 20 10 -10
149 17 11 -6
150 11 11 0
151 11 11 0
152 15 11 -4
153 17 10 -7
154 11 7 -4
155 20 9 -11
156 22 9 -13
157 5 12 7
158 19 11 -8
159 17 9 -8
160 14 9 -5
161 8 5 -3
162 11 6 -5
163 13 9 -4
164 14 6 -8
165 5 11 6
166 7 8 1
167 6 5 -1
168 19 11 -8
169 10 20 10
170 22 5 -17
171 14 11 -3
172 10 11 1
173 18 18 0
174 19 13 -6
175 13 17 4
176 17 18 1
177 9 11 2
178 19 12 -7
179 17 12 -5
180 12 14 2
181 21 19 -2
182 18 8 -10
183 6 11 5
184 12 13 1
185 21 7 -14
186 14 7 -7
187 7 5 -2
188 5 9 4
189 16 13 -3
190 20 6 -14
191 9 14 5
192 12 6 -6
193 2 5 3
194 7 NA NA
195 9 7 -2
196 5 8 3
197 13 9 -4
198 14 11 -3
199 7 9 2
200 15 15 0
201 12 17 5
202 20 15 -5
203 19 11 -8
204 17 22 5
205 12 4 -8
206 11 5 -6
207 17 7 -10
208 18 10 -8
209 5 7 2
211 23 17 -6
212 4 8 4
214 24 15 -9
215 6 14 8
216 6 8 2
217 12 10 -2
218 17 10 -7
219 9 11 2
220 11 14 3
221 16 10 -6
222 17 11 -6
223 17 11 -6
224 11 12 1
225 21 15 -6
226 17 14 -3
228 6 10 4
229 17 12 -5
230 17 11 -6
231 9 6 -3
235 19 19 0
236 7 5 -2
237 14 9 -5
238 7 3 -4
239 14 7 -7
240 9 9 0
241 12 11 -1

3.2. Hierarchical Modelling of Species Communities

All variables

Prediction of specific richness:

Difference between predicted and observed taxa richness
station_id observed predicted difference
127 1 6 5
128 12 4 -8
129 15 2 -13
130 4 6 2
131 10 3 -7
132 10 3 -7
134 9 5 -4
135 8 7 -1
136 9 9 0
137 11 9 -2
138 12 11 -1
139 13 4 -9
140 14 6 -8
141 14 9 -5
142 22 5 -17
143 12 5 -7
144 12 6 -6
145 11 5 -6
146 10 6 -4
147 9 7 -2
148 20 10 -10
149 17 9 -8
150 11 5 -6
151 11 11 0
152 15 9 -6
153 17 4 -13
154 11 9 -2
155 20 10 -10
156 22 7 -15
157 5 4 -1
158 19 4 -15
159 17 8 -9
160 14 2 -12
161 8 2 -6
162 11 3 -8
163 13 1 -12
164 14 3 -11
165 5 4 -1
166 7 3 -4
167 6 1 -5
168 19 3 -16
169 10 11 1
170 22 5 -17
171 14 5 -9
172 10 13 3
173 18 4 -14
174 19 5 -14
175 13 3 -10
176 17 5 -12
177 9 7 -2
178 19 8 -11
179 17 4 -13
180 12 8 -4
181 21 5 -16
182 18 5 -13
183 6 6 0
184 12 8 -4
185 21 6 -15
186 14 4 -10
187 7 1 -6
188 5 8 3
189 16 11 -5
190 20 3 -17
191 9 4 -5
192 12 3 -9
193 2 7 5
194 7 NA NA
195 9 5 -4
196 5 2 -3
197 13 3 -10
198 14 6 -8
199 7 15 8
200 15 15 0
201 12 13 1
202 20 4 -16
203 19 3 -16
204 17 4 -13
205 12 4 -8
206 11 4 -7
207 17 6 -11
208 18 6 -12
209 5 4 -1
211 23 9 -14
212 4 12 8
214 24 13 -11
215 6 7 1
216 6 0 -6
217 12 4 -8
218 17 5 -12
219 9 0 -9
220 11 3 -8
221 16 9 -7
222 17 8 -9
223 17 9 -8
224 11 7 -4
225 21 9 -12
226 17 9 -8
228 6 1 -5
229 17 4 -13
230 17 4 -13
231 9 4 -5
235 19 6 -13
236 7 10 3
237 14 5 -9
238 7 3 -4
239 14 11 -3
240 9 12 3
241 12 6 -6
Abiotic parameters

Prediction of specific richness:

Difference between predicted and observed taxa richness
station_id observed predicted difference
127 1 2 1
128 12 3 -9
129 15 3 -12
130 4 3 -1
131 10 3 -7
132 10 4 -6
134 9 7 -2
135 8 7 -1
136 9 8 -1
137 11 8 -3
138 12 9 -3
139 13 3 -10
140 14 5 -9
141 14 6 -8
142 22 7 -15
143 12 7 -5
144 12 7 -5
145 11 9 -2
146 10 10 0
147 9 10 1
148 20 10 -10
149 17 7 -10
150 11 4 -7
151 11 9 -2
152 15 7 -8
153 17 3 -14
154 11 8 -3
155 20 4 -16
156 22 6 -16
157 5 3 -2
158 19 3 -16
159 17 7 -10
160 14 3 -11
161 8 5 -3
162 11 6 -5
163 13 4 -9
164 14 8 -6
165 5 10 5
166 7 4 -3
167 6 4 -2
168 19 2 -17
169 10 11 1
170 22 5 -17
171 14 3 -11
172 10 6 -4
173 18 6 -12
174 19 5 -14
175 13 5 -8
176 17 5 -12
177 9 7 -2
178 19 10 -9
179 17 6 -11
180 12 11 -1
181 21 7 -14
182 18 5 -13
183 6 7 1
184 12 7 -5
185 21 5 -16
186 14 5 -9
187 7 4 -3
188 5 12 7
189 16 5 -11
190 20 3 -17
191 9 3 -6
192 12 2 -10
193 2 7 5
194 7 NA NA
195 9 5 -4
196 5 2 -3
197 13 2 -11
198 14 7 -7
199 7 13 6
200 15 10 -5
201 12 9 -3
202 20 3 -17
203 19 4 -15
204 17 5 -12
205 12 4 -8
206 11 6 -5
207 17 7 -10
208 18 8 -10
209 5 3 -2
211 23 5 -18
212 4 8 4
214 24 6 -18
215 6 5 -1
216 6 0 -6
217 12 3 -9
218 17 3 -14
219 9 0 -9
220 11 2 -9
221 16 10 -6
222 17 12 -5
223 17 9 -8
224 11 7 -4
225 21 13 -8
226 17 8 -9
228 6 5 -1
229 17 4 -13
230 17 3 -14
231 9 5 -4
235 19 8 -11
236 7 7 0
237 14 3 -11
238 7 3 -4
239 14 9 -5
240 9 11 2
241 12 4 -8
Exposure indices

Prediction of specific richness:

Difference between predicted and observed taxa richness
station_id observed predicted difference
127 1 5 4
128 12 3 -9
129 15 5 -10
130 4 3 -1
131 10 3 -7
132 10 4 -6
134 9 3 -6
135 8 4 -4
136 9 3 -6
137 11 5 -6
138 12 6 -6
139 13 3 -10
140 14 3 -11
141 14 2 -12
142 22 1 -21
143 12 2 -10
144 12 2 -10
145 11 2 -9
146 10 2 -8
147 9 4 -5
148 20 9 -11
149 17 6 -11
150 11 6 -5
151 11 9 -2
152 15 9 -6
153 17 6 -11
154 11 10 -1
155 20 7 -13
156 22 5 -17
157 5 3 -2
158 19 4 -15
159 17 8 -9
160 14 2 -12
161 8 7 -1
162 11 7 -4
163 13 2 -11
164 14 7 -7
165 5 8 3
166 7 3 -4
167 6 3 -3
168 19 4 -15
169 10 9 -1
170 22 3 -19
171 14 5 -9
172 10 3 -7
173 18 4 -14
174 19 7 -12
175 13 1 -12
176 17 4 -13
177 9 4 -5
178 19 3 -16
179 17 3 -14
180 12 6 -6
181 21 4 -17
182 18 3 -15
183 6 3 -3
184 12 3 -9
185 21 4 -17
186 14 2 -12
187 7 2 -5
188 5 4 -1
189 16 8 -8
190 20 3 -17
191 9 7 -2
192 12 3 -9
193 2 10 8
194 7 NA NA
195 9 3 -6
196 5 3 -2
197 13 4 -9
198 14 3 -11
199 7 12 5
200 15 11 -4
201 12 11 -1
202 20 3 -17
203 19 5 -14
204 17 2 -15
205 12 3 -9
206 11 3 -8
207 17 6 -11
208 18 4 -14
209 5 4 -1
211 23 4 -19
212 4 11 7
214 24 9 -15
215 6 3 -3
216 6 5 -1
217 12 3 -9
218 17 4 -13
219 9 3 -6
220 11 3 -8
221 16 5 -11
222 17 9 -8
223 17 9 -8
224 11 5 -6
225 21 9 -12
226 17 8 -9
228 6 7 1
229 17 3 -14
230 17 3 -14
231 9 1 -8
235 19 2 -17
236 7 8 1
237 14 4 -10
238 7 4 -3
239 14 10 -4
240 9 10 1
241 12 9 -3

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